VICTORIA. Otway Range, Triple Falls, Phillips Track, hypogeal in mixed eucalyptus forest, 22 Sept. 1982, K & G. Beaton 57.
NEW ZEALAND. Wellington, Chateau Tongariro, April 1931, J. Carter ( type of G. novaezelandiae, K); Horopito, Mangatorutoru Stream, March 1948, J. M. Dingley.
U.S.A. Oregon, Benton County, 14 Nov. 1924, S. M Zeller 2260, ex herb. Hawker.

Gasterocarp 0.5-3 cm diam., subglobose, pyriform to ellipsoid or irregular, lobed, with basal attachment. Peridium whitish to ochraceous, with a distinctly greenish yellow tint, not cyanescent on bruising, smooth and glabrous. Gleba pale brown drying to dark brown, firm, loculate, consisting of irregular, partially filled, minute chambers. Tramal plates variable, mostly about 100µm thick, consisting of a broad, hyaline hymenophoral trama and very narrow subhymenial layers; peridial context well developed, forming a thick layer, similar in structure to the hymenophoral trama; clamp-connexions absent. Columella very reduced, occasionally forming very short radiating branches; sterile base present but small and inconspicuous. Spores 10.5-15 x 6-7.5 (12 ±0.5 x 7 ± 0.5) µm (excl. costae), Q = 1.71; symmetric, ellipsoid to short fusoid, sometimes with a papillate apex, deep rusty brown, with a thickened wall, smooth but forming four or five longitudinal costae, overlaid by a hyaline myxosporium which sometimes becomes detached at the base. Basidia 15-20 x 7.5-9µm, clavate to cylindric, bearing four, sometimes two, sterigmata. Hymenophoral trama regular, hyaline, non-gelatinized, of parallel, thinwalled hyphae, 2-10 µm diam. Subhymenial layer very narrow, interwoven. Peridiopellis a repent epicutis, 50-75 µm thick, of subparallel or partial woven hyaline, thin-walled hyphae, 3-10 µm diam.

This species has been described from New Zealand under two separate names, Hymenogaster pachydermis Zeller & Dodge and Gautieria novaezelandiae G. H. Cunn. Cunningham (1944: 67) listed H. pachydermis as a synonym under Hysterangium schlerodermun (Cooke) G. H. Cunn., but examination of the authentic collection by Zeller reveals it to have the pigmented, costate spores of a Chamonixia species as originally described. Chamonixia pachydermis differs from C. mucosa in the non-cyanescent, greenish peridium and the more deeply coloured spores.

Type: Mycorrhizal Fungi; Description: Basidiomata epigeous or subepigeous, subglobose to pyriform, ochraceous to bright greenish brown, up to 25 mm in diameter; peridium composed of a single white to cream pseudoparenchymatous layer. Gleba chocolate brown, compact; columella reduced to a few tenuous branches. Basidiospores elliptical, 0-septate, 10–15 × 8–10 μm, with 4–5 longitudinal ribs, golden brown.

Distribution: Auckland, Coromandel, Waikato, Bay of Plenty, Taupo, Wellington, Nelson, Gisborne, Buller, Westland, Fiordland, Marlborough Sounds, North Canterbury, Mid Canterbury, Otago Lakes, Southland.; 1st Record: Chu-Chou & Grace (1983b: as Gauteria novaezelandiae).

Significance: Recorded as a close associate of Nothofagus spp. (Chu-Chou & Grace 1983b).; Host(s): Nothofagus fusca, N. menziesii, N. solandri var. solandri, N. solandri var. cliffortioides, N. truncata.