Capitula small to large, solitary, or infl. variously compound; receptacle alveolate, flat or convex; phyll. imbricate in several series, margins dry or scarious. Ray-florets, if present, ligulate to filiform, pistillate; disk-florets perfect, tubular. Anther-cells us. acute at base; style-arms of disk-florets flattened, with short obtuse or acute unreceptive tips. Achenes ribbed or striate, us. pubescent to pilose; pappus of unequal barbellate bristles, often slightly thickened at tips. Shrubs or trees with alt. or opp. lvs, these mainly tomentose below. About 130 spp., mainly of N.Z. and Australia, with a few in Lord Howe Id and New Guinea; the N.Z. spp. endemic.