Capitula small, solitary or clustered (occ. only 2 together), sessile or on short scapes, heterogamous, discoid; phyll. few, in 2 series, subequal; receptacle nude. Anthers obtuse to pointed at base; style-arms of disk-florets short, truncate; pappus 0. Achenes 4-angled to terete or fusiform. Some 20 spp. of small ± mosslike glab. (or very nearly so) herbs with alt. entire lvs; of Fuegia, Falkland Is, Australia, Tasmania, N.Z. (these endemic).

Perennial herbs, often very small, low, mat- or cushion-forming or caespitose, rarely up to 12 cm high. Leaves rosulate, alternate, often densely imbricate, sometimes a few cauline or bract-like, linear-oblanceolate or oblong-ovate, glabrous or very rarely with septate hairs, rarely keeled or flat but still somewhat crassulate-coriaceous, with sunken multicellular glands; apex retuse, obtuse or apiculate; margins entire, sometimes scariose, often with septate hairs towards the base; petiole sheath-like or rarely a true sheath, often purple; stomata adaxial and abaxial. Capitula disciform, heterogamous, solitary and terminal or seldom 2-10 corymbosely arranged, up to 5 mm wide, sessile to pedunculate; peduncle often elongated in fruit, glabrous or rarely septate hairy; bracts leaf-like, phyllary-like, or absent. Phyllaries (3-) 5-14 (-19), ± imbricate, free, equal, in 1-3 series, oblong-linear or obovate, often with microscopic glands at the base, sometimes with translucent ducts; nerves 1 or rarely 5 or absent; apex obtuse-acute or retuse; margins entire or seldom denticulate, ± scariose, often tinged purple. Receptacle naked and flat. Outer florets female, fertile, tubular or cyathiform, in one series, glabrous; corolla 3-4 (-5)-lobed, basally often bulbous, white, purple, pale green, or yellowish, often two-coloured; vascular tissue in lobe margin, central, or absent; style glabrous, bilobed; style-branches short, ovate. Central florets functionally male, campanulate, with an undivided, truncate style lacking sweeping hairs and stigmatic papillae, or central florets perfect, ± tubular to slightly campanulate, with a divided, truncate style having short, acute sweeping hairs and stigmatic papillae; corolla 4-lobed but occasionally 2-, 3- to 5-lobed, white, purple, pale green or yellowish, often two-coloured, glabrous; vascular tissue in lobe margin, central, or both central and marginal, or absent; anthers caudate, 0.5-1.5 mm long; appendage oblong; endothecium polarized orrarely partly radial. Cypsela straight or rarely slightly curved, ellipsoid-obvoid or somewhat terete, nerved or ribbed, glabrous, minutely papillose or hairy; apex smooth, rimmed, crowned or with small horns or awns, rarely beaked. Pappus absent.