Leptinella dioica Hook.f.
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Leptinella dioica Hook.f., Bot. Antarct. Voy. II. (Fl. Nov.-Zel.) Part I, 129 (1852)
Leptinella dioica Hook.f.
Nomenclature
Hook.f.
Hook.f.
1852
129
ICN
Leptinella dioica Hook.f.
species
Leptinella dioica
Classification
Vernacular names
Synonyms
Descriptions
Stems rather stout, much-branched, creeping, rooting, glab. or with scattered hairs; plants forming dense to rather open patches up to c. 3 dm. diam. Lvs tufted at nodes or alt. Lamina membr., ± flaccid, dull green, glab. or nearly so, broad- to narrow-obovate to spathulate, obtuse, ± 20-30 × 5-9 mm. (but see also description of vars) on slender petioles up to ± 15 mm. long; upper portion crenately toothed, lower portion pinnatifid to pinnatisect; segs 6-8 pairs, subopp., oblong, subacute, entire, or upper margins with a few teeth, lowest segs minute. Scapes axillary, ± pubescent, c. 3 cm. long, nude; receptacle convex. Capitula of ♀ 8-10 mm. diam.; phyll. several in 3-4 series, inturned and concealing florets. Capitula of ♂ 4-7 mm. diam.; phyll. few. in c. 2 series, orbicular-oblong, pubescent, with broad purple scarious margins. Florets eglandular; ♀ subconic, inflated at base, minutely 4-toothed at apex; ♂ funnelform. Achenes obovoid, slightly curved, rounded at back, c. 2 mm. long, glab.
Leptinella dioica Hook.f.
2n = 260
A creeping, fleshy perennial herb. Rhizomes at or near soil surface, green or dark, flexible and ± pilose, becoming pale, wiry and glabrous if buried; branches uncommon, us. single at flowering nodes; leaves in two rows, single at the apex, 0.3-3.0 cm apart. Short shoots alternate on both sides of the rhizome, with up to 5 clustered leaves, rarely converted into rhizomes with distant leaves. Roots us. slender and weak, up to 0.8 mm diam. Leaves very variable in size, shape, and divisions, simple to incised-pinnatifid, occ. pinnatifid, 0.7-12.0 × 0.3-1.5 cm; blade 0.5-7.0 cm, narrowly to broadly obovate or elliptic, fleshy, green, or glaucous, us. without dark pigment, ± glabrous but dotted with sunken glandular hairs, midrib not raised on vertical surface, lobes (pinnae or teeth) 4-12 pairs, distant or the distal ones overlapping, oblong to orbicular; proximal lobes cut to rhachis, sinuses of distal lobes usually not reaching rhachis, sts cut only 1/5 to rhachis at widest part of the leaf; teeth often absent but up to 6 per lobe, on the distal and outer margins, small, triangular, obtuse or rounded, apiculate. Peduncles us. borne on rhizomes, ca. equal to leaves, 1-6 cm, nude or with 1 simple bract, sparsely villous. Us, dioecious, occ. monoecious. Pistillate heads 2-7 mm diam.. up to 10 mm in fruit; surface convex; involucre urceolate; phyllaries 10-30 in 3 or more unequal rows, broadly elliptic, green, glabrous, with a wide, often brown-tipped, scarious margin; inner phyllaries grow after anthesis to enclose subglobose fruiting head; florets 10-80, ca. 2-5 mm long, curved, yellow-green; corolla slightly longer than wide, with unequal teeth. Staminate heads 3-8 mm diam.; involucre hemispherical; phyllaries 5-10 in 1-2 subequal rows, not growing after anthesis; florets slightly more numerous. Bisexual heads predominantly staminate. Achenes up to 1.9 × 1.0 mm, slightly compressed, almost round or irregularly angled, with a pale unwrinkled papery surface turning brown and smooth. Flowers in spring and summer.
Leaves incised-pinnatifid (pinnatifid from Dunedin south), often ± glaucous, very variable in shape and division, narrowly to broadly elliptic or obovate; lobes 4-12 pairs, oblong to suborbicular, often toothed; lowermost sinuses on each side extending to rhachis; base of most proximal lobes ± truncate; terminal lobe not distinctly wider (between sinuses) than lateral lobes. Dioecious, heads on one plant either staminate or pistillate only; inner phyllaries of pistillate heads elongating after anthesis, enclosing fruit. Pollen grains with even spines and walls.
Leaves yellow-green, rhomboid, obovate or elliptic, sts entire but us. incised-pinnatifid with 1-5 triangular entire lobes or teeth on each side; lowermost sinus on each side not usually extending to rhachis; base of blade cuneate; terminal lobe distinctly wider than lateral lobes. Monoecious; heads on each plant a mixture of staminate heads and bisexual heads with us. 1 or 2 but sts up to 5 pistillate florets in 1 incomplete row surrounding 10-30 staminate florets. Pollen grains functional but with walls irregular in thickness and spines lacking or underdeveloped. Chromosome number not known. Figs. 3, 4.
Folia interdum integra sed plerumque incisa-pinnatiflda cum 1-5 triangularibus, integris lobis utrimque; loba terminalis lobis lateralibus maior. Monoica; capitula cum 0-5 femineis flosculis 10-30 masculos flosculos cingentibus.
Taxonomic concepts
Cotula dioica (Hook.f.) Hook.f.
Leptinella dioica Hook.f.
Cotula dioica (Hook.f.) Hook.f.
Leptinella dioica Hook.f.
Cotula dioica (Hook.f.) Hook.f. subsp. dioica
Cotula dioica (Hook.f.) Hook.f. subsp. dioica
Cotula dioica (Hook.f.) Hook.f. subsp. dioica
Cotula dioica subsp. monoica D.G.Lloyd
Cotula dioica subsp. monoica D.G.Lloyd
Cotula dioica var. crenatifolia Kirk
Leptinella dioica Hook.f.
Cotula dioica var. crenatifolia Kirk
Leptinella dioica Hook.f.
Cotula dioica var. obscura (Kirk) Cheeseman
Cotula dioica var. obscura (Kirk) Cheeseman
Cotula obscura Kirk
Cotula obscura Kirk
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Leptinella dioica Hook.f.
Collections
Identification keys
Metadata
29ffdfdf-67e1-472b-a44b-c3886a8474c1
scientific name
Names_Plants
1 January 2000
8 November 2005