Leaves incised-pinnatifid (pinnatifid from Dunedin south), often ± glaucous, very variable in shape and division, narrowly to broadly elliptic or obovate; lobes 4-12 pairs, oblong to suborbicular, often toothed; lowermost sinuses on each side extending to rhachis; base of most proximal lobes ± truncate; terminal lobe not distinctly wider (between sinuses) than lateral lobes. Dioecious, heads on one plant either staminate or pistillate only; inner phyllaries of pistillate heads elongating after anthesis, enclosing fruit. Pollen grains with even spines and walls.