Pileus to 3 mm diam., convex to inverted saucer-shaped, nutant, hymenium rarely exposed upwards, convex, white, pruinose to minutely setose, dry. Lamellae none, hymenophore smooth, white. Pseudostipe up to 6 X up to 0.4 mm, central, inserted and gradually confluent into apex (centre) of pileus, cylindrical or slightly enlarged towards apex, whitish to glassy-hyaline, rather fragile, minutely pruinose overall, at base minutely strigose-hairy, insititious (basal disc or byssus absent), dry, solid, solitary in groups. Odour absent. Basidiospores 14-19 X 6.5-7.5 µm, elliptic to subfusiform, inequilateral in profile, smooth, hyaline, inamyloid. Basidia 30-40 X 11-13 µm, tetrasporic, cylindric to subclavate, sterimata up to 10 µm long, clamped. Hymenial cystidia scattered, 60-120 X 10-15 µm, slender fusiform with subglobose apex (6-10 µm diam.), thin-walled, hyaline, often filled with strongly refringent cell sap. Pileipellis hymeniform, composed of clavate or vesiculose cells 15-45 X 18-32 µm, non-gelatinous, thin-walled, hyaline. Pilocystidia scattered, 35-100 X 5-12 µm, subcylindric to slender fusiform, apex rounded or slightly swollen, thin-walled, hyaline. Oleiferous hyphae absent. Stipe tissue monomitic. Stipe cortical hyphae (4-8 µm diam., cylindric, parallel, thin walled, hyaline. Caulocystidia inconspicuous, rarely differentiated, 6-50 X 6-11 µm, mostly as blunt and broad projections, thin-walled, hyaline, clamped.

On rotting leaves of Phormium tenax J.R. & G. Forst. (Phormiaceae). Known from South Island of New Zealand.
Commentary

Pileus usque ad 3 mm latus., inverso-convexus vel -cupuliformis, pendulans, albus, pruinosus vel tomentosus, siccus. Lamellae nullae, hymenium glabrum, album. Stipes usque ad 6 X 0.4 mm, centraliter ad apicem affixus, cylindricus, albidus vel hyalinus, fragilis, minute pruinosus, ad basim strigoso. Inodorus. Basidiosporae 14-19 X 6.5-7.5 µm, ellipticae vel subfusiformes, leves, hyalinae, inamyloideae. Basidia 30-40 X 11-13 µm, tetrasporica, fibulata. Cystidia hymeniales rara, 60-120 X 10-15 µm, fusiformia vel sublecythiformia, membrana tenuitunicata hyalineque instructa. Pileipellis hymeniformis, e cellulis clavatis vel vesiculosis (15-45 X 18-32 µm). Pilocystidia rara, 35-100 X 5-12 µm, subcylindrica vel fusiformia. Caulocystidia inconspicua. Ad folia putrida Phormii tenacis, Novazelandia. PDD, holotypus.

The new genus Anastrophella is characterised by nutant basidiomes with inverted-cupulate pilei and smooth hymenophore, a relatively long, central pseudostipe, and a hymeniform pileipellis of smooth, vesiculose cells and pilocystidia. It is allied with the genera Gloiocephala Massee (1892) and Physalacria Peck (1882). In essence, Anastrophella is a Gloiocephala with an upside down pileus and correspondingly nutant habit.
Singer (1976: 311) erected the monotypic taxon Physalacria sect. Pileolina Singer for P. subpeltata Redhead [a binomial that wasn't published validly until 1979 (Redhead 1979)], separating the species from Physalacria s. str. (Berthier 1985) because of the discoid-pileate, pseudostipitate basidiomes. Subsequently, Singer (1986: 374) transferred Physalacria sect. Pileolina and Gloiocephala sect. Sessiles Singer (1976) to the genus Deigloria Agerer (1980), thereby restricting Physalacria to those species with stalked-capitate basidiomes, i.e. with a globose to obclavate, hollow pileus that is fertile over the entire external surface and sits atop a short pseudostipe. We concur with Agerer (1980) that Ph. subpeltata is distinct from Deigloria, differing in forming a well-developed pseudostipe, a discoid-nutant pileus, a pileipellis composed of clavate to vesiculose cells lacking apical appendages and with fusiform-subcapitate pilocystidia and capitate gloeocystidia. In contrast, Deigloria is characterised by sessile-cupulate basidiomes, a non-fertile surface composed of mucronate or broom-like elements, and dermatogloeocystidia with an apical whorl of finger-like appendages (Agerer 1980).
We agree with Singer (1986) that the species he placed previously in Gloiocephala sect. Sessiles rightfully belong in Deigloria, but we consider his sect. Pileolina as distinct from both Physalacria and Deigloria. Accordingly we propose the new genus Anastrophella to accommodate Ph. subpeltata and the New Zealand specimen. For further discussion about taxonomic relationships, cf. Hispidocalyptella.

Holotype
New Zealand, South Island, Westland, Rotomanu, Lady Lake, 25.iii. 1983, Horak (PDD; ZT 2100, isotype).