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Horak, E.; Desjardin, D.E. 1994: Reduced marasmioid and mycenoid agarics from Australasia. Australian Systematic Botany 7: 153-170.

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Horak, E.; Desjardin, D.E. 1994: Reduced marasmioid and mycenoid agarics from Australasia. Australian Systematic Botany 7: 153-170.
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Pileus minutus, nutans, inverso-discoideus vel inverso-cupulatus, hymenio interiori instructus, pruinosus, albus. Hymenium leve. Pseudostipes centralis, bene evolutus, pruinosus, insititius. Basidiosporae ellipsoideae vel subfusiformia, leves, hyalinae, inamyloideae. Cystidia hymeniales praesentes vel nulla, capitata. Pileipellis hymeniformis e cellulis vesiculosis levibusque et raris pilocystidiis ventricosis vel lageniformibus et gloeocystidiis elongato-capitatis.
Etymology
Anastrepho, turned upside down, inverted: referring to the alignment of the pileus with respect to the stipe.
Physalacria subpeltata Redhead. Mycotaxon 10 (1): 46. 1979.br Anastrophella subpeltata (Redhead) Horak & Desjardin, comb. nov.
Pileus to 3 mm diam., convex to inverted saucer-shaped, nutant, hymenium rarely exposed upwards, convex, white, pruinose to minutely setose, dry. Lamellae none, hymenophore smooth, white. Pseudostipe up to 6 X up to 0.4 mm, central, inserted and gradually confluent into apex (centre) of pileus, cylindrical or slightly enlarged towards apex, whitish to glassy-hyaline, rather fragile, minutely pruinose overall, at base minutely strigose-hairy, insititious (basal disc or byssus absent), dry, solid, solitary in groups. Odour absent. Basidiospores 14-19 X 6.5-7.5 µm, elliptic to subfusiform, inequilateral in profile, smooth, hyaline, inamyloid. Basidia 30-40 X 11-13 µm, tetrasporic, cylindric to subclavate, sterimata up to 10 µm long, clamped. Hymenial cystidia scattered, 60-120 X 10-15 µm, slender fusiform with subglobose apex (6-10 µm diam.), thin-walled, hyaline, often filled with strongly refringent cell sap. Pileipellis hymeniform, composed of clavate or vesiculose cells 15-45 X 18-32 µm, non-gelatinous, thin-walled, hyaline. Pilocystidia scattered, 35-100 X 5-12 µm, subcylindric to slender fusiform, apex rounded or slightly swollen, thin-walled, hyaline. Oleiferous hyphae absent. Stipe tissue monomitic. Stipe cortical hyphae (4-8 µm diam., cylindric, parallel, thin walled, hyaline. Caulocystidia inconspicuous, rarely differentiated, 6-50 X 6-11 µm, mostly as blunt and broad projections, thin-walled, hyaline, clamped.
On rotting leaves of Phormium tenax J.R. & G. Forst. (Phormiaceae). Known from South Island of New Zealand.
Commentary
Pileus usque ad 3 mm latus., inverso-convexus vel -cupuliformis, pendulans, albus, pruinosus vel tomentosus, siccus. Lamellae nullae, hymenium glabrum, album. Stipes usque ad 6 X 0.4 mm, centraliter ad apicem affixus, cylindricus, albidus vel hyalinus, fragilis, minute pruinosus, ad basim strigoso. Inodorus. Basidiosporae 14-19 X 6.5-7.5 µm, ellipticae vel subfusiformes, leves, hyalinae, inamyloideae. Basidia 30-40 X 11-13 µm, tetrasporica, fibulata. Cystidia hymeniales rara, 60-120 X 10-15 µm, fusiformia vel sublecythiformia, membrana tenuitunicata hyalineque instructa. Pileipellis hymeniformis, e cellulis clavatis vel vesiculosis (15-45 X 18-32 µm). Pilocystidia rara, 35-100 X 5-12 µm, subcylindrica vel fusiformia. Caulocystidia inconspicua. Ad folia putrida Phormii tenacis, Novazelandia. PDD, holotypus.
The new genus Anastrophella is characterised by nutant basidiomes with inverted-cupulate pilei and smooth hymenophore, a relatively long, central pseudostipe, and a hymeniform pileipellis of smooth, vesiculose cells and pilocystidia. It is allied with the genera Gloiocephala Massee (1892) and Physalacria Peck (1882). In essence, Anastrophella is a Gloiocephala with an upside down pileus and correspondingly nutant habit.
Singer (1976: 311) erected the monotypic taxon Physalacria sect. Pileolina Singer for P. subpeltata Redhead [a binomial that wasn't published validly until 1979 (Redhead 1979)], separating the species from Physalacria s. str. (Berthier 1985) because of the discoid-pileate, pseudostipitate basidiomes. Subsequently, Singer (1986: 374) transferred Physalacria sect. Pileolina and Gloiocephala sect. Sessiles Singer (1976) to the genus Deigloria Agerer (1980), thereby restricting Physalacria to those species with stalked-capitate basidiomes, i.e. with a globose to obclavate, hollow pileus that is fertile over the entire external surface and sits atop a short pseudostipe. We concur with Agerer (1980) that Ph. subpeltata is distinct from Deigloria, differing in forming a well-developed pseudostipe, a discoid-nutant pileus, a pileipellis composed of clavate to vesiculose cells lacking apical appendages and with fusiform-subcapitate pilocystidia and capitate gloeocystidia. In contrast, Deigloria is characterised by sessile-cupulate basidiomes, a non-fertile surface composed of mucronate or broom-like elements, and dermatogloeocystidia with an apical whorl of finger-like appendages (Agerer 1980).
We agree with Singer (1986) that the species he placed previously in Gloiocephala sect. Sessiles rightfully belong in Deigloria, but we consider his sect. Pileolina as distinct from both Physalacria and Deigloria. Accordingly we propose the new genus Anastrophella to accommodate Ph. subpeltata and the New Zealand specimen. For further discussion about taxonomic relationships, cf. Hispidocalyptella.
Holotype
New Zealand, South Island, Westland, Rotomanu, Lady Lake, 25.iii. 1983, Horak (PDD; ZT 2100, isotype).
Pileus 2-7 mm diam., at first convex, soon becoming plane or depressed at centre with up turned margin, white, cream or pale yellow, pruinose to velvety (minutely hairy-strigose under hand-lens), indistinctly sulcate towards margin, gelatinous in fresh condition (pileipellis separable). Lamellae 2-6, venose to fold-like, attached to apex of stipe, occasionally hymenophore smooth, cream, edges concolorous. Stipe 3-7 X up to I mm, always central, cylindric, equal, whitish to cream, minutely pruinose overall, dry, solid, solitary in groups, insititious (basal disc or byssus absent). Basidiospores 16-19 X 8-9.5 µm, broadly elliptic, slightly inequilateral in profile, smooth, hyaline, inamyloid. Basidia 40-45 X 11-15 µm, tetrasporic, subcylindric to clavate, sterigmata up to 9 µm long, clamped. Cheilo- and pleuro-cystidia numerous, 60-115 X 20-30 µm, fusiform to lageniform, apex rounded or indistinctly capitate, thin-walled, smooth, hyaline. Pileipellis hymeniform, composed of vesiculose to clavate smooth cells 20-30 X 10-20 µm, subgelatinous, hyaline, thin-walled. Pilocystidia numerous, 40-85 X 8-12 µm, fusiform with long, gradually tapering neck, apex rounded or sub acute, thin-walled, hyaline. Oleiferous hyphae absent. Stipe tissue monomitic. Stipe cortical hyphae cylindric, 5-10 pm diam., smooth, thin-walled. Caulocystidia numerous. 20-110 X 3-7 µm, trichiform or subfusiform with gradually tapering apex, tips rounded or pointed, thin-walled, hyaline.
On rotting leaves of Phormium tenax J.R. & G. Forst. (Phormiaceae). Known from South Island of New Zealand.
Pileus 2-7 mm latus plano-convexus dein depressus, albus vel pallide stramineus, minute pruinosus, subgelatinosus. Lamellae nonnullae (2-6), venosae, rariter desunt, pallide stramineae. Stipes 3-7 X up to 1 mm, centralis, cylindricus, pileo concolor, minute pruinosus. Inodorus. Basidiosporae 16-19 X 8-9.5 µm, ellipticae, leves, hyalinae, inamyloideae. Basidia 40-45 X 11-15 µm. tetrasporica, fibulata. Cheilo- et pleuro-cystidia 60-115 X 20-30 µm, fusiformia, subcapitata, tenuitunicata, hyalina. Pileipellis hymeniformis e cellulis vesiculoso-clavatis (20-30 X 10-20 µm). Pilocystidia numerosa, 40-85 X 8-12 µm, fusiformia, apicaliter attenuata, tenuitunicata, hyalina. Caulocystidia 20-110 X 3-7 µm, trichiformia vel subfusiformia. Ad folia putrida Phormii tenacis. Novazelandiae. PDD, holotypus
Gloiocephala phormiorum belongs in sect. Gloiocephala subsect. Macrosporae Singer (1976) where its closest ally appears to be G. longispenna Singer (from Brazil: Singer 1976). The New Zealand taxon is distinguished by relatively large basidiomes with 2-6 fold-like lamellae and a well-developed, white, central stipe: broad basidiospores (8-9.5 µm diam.); fusiform to lageniform, thin-walled hymenial cystidia 60-110 µm long, and fusiform-rostrate pilocystidia 40-85 µm long. In comparison, G. longisperma forms small basidiomes with a more reduced hymenophore and black stipe base, narrower basidiospores (3-4 µm), ventricose hymenial cystidia 30-40 µm long, and pilocystidia 100-200 µm long (fide Singer 1976). The present new species differs from others in the subsection by lacking metuloid hymenial cystidia, having much larger basidiospores, and in its fructification on Phormiaceae.
Holotype
New Zealand, Canterbury, Ashley, Mt Grey, Kowai River Reserve, 22.ix.1967, Horak (PDD; ZT 67-127, isotype).
Pileus to 4 mm diam., hemispherical, becoming convex to flat expanded, whitish, minutely pruinose to velutinous (under hand-lens), dry, tough. Lamellae none, hymenophore smooth or with few venose radial folds. Stipe to 8 X 0.3 mm, central to slightly eccentric, cylindrical, slender, horse-hair-like, apex whitish, dark brown to black towards base: apex pruinose, becoming smooth downwards, dry, elastic, solid, solitary in groups, insititious (basal disc or byssus absent). Odour absent. Basidiospores 10-1 I X 5-5.5 µm, broadly fusoid to subamygdaliform, inequilateral in profile, smooth, hyaline, inamyloid. Basidia 25-35 X 8-9 µm, tetrasporic, subclavate (to cylindric), sterigmata up to 4 µm long, clamped. Hymenial cystidia absent or rarely encountered towards the margin, shape and size as caulocystidia. Pileipellis hymeniform, composed of slender clavate or vesiculose, non-diverticulate cells 20-25 X 8-18 µm, thick-walled (1-3 µm) especially at apex, hyaline, often encrusted with orange-brown pigment (in KOH). Pilocystidia numerous, 30-55 X 4-8 µm, slender fusoid with capitate apex (3-6 µm diam.), thick-walled except at globose apex, hyaline, tips occasionally bearing a resinous drop. Oleiferous hyphae absent. Stipe tissue monomitic. Stipe cortical hyphae 3-8 µm diam., parallel, smooth, non-gelatinous, hyaline, inamyloid, clamped. Caulocystidia numerous, 30-45 X 5-9 µm, shape like pilocystidia or typically tibiiform to lecythiform, globose apex 6-8 µm diam., often with resinous cap, thin-walled (< 1.5 µm diam.), hyaline.
On rotting leaves of Phormium tenax J.R. & G. Forst. (Phormiaceae). Known from North Island of New Zealand.
Pileus usque ad 4 mm latus, hemisphaerical dein convexo-applanatus, albidus, minute pruinosus vel tomentosus, siccus. Lamellae nullae, hymenium glabrum vel subplicatum. Stipes usque ad 8 X 0.3 mm, centralis vel subeccentricus, cylindricus, tenax, apaliter albidus, basin versus fuscus vel niger, apicaliter pruinosus, deorsum glabrus, siccus. Inodorus. Basidiosporae 10-11 X 5-5.5 µm, fusoideae vel subamygdaliform, leves, hyalinae, inamyloideae. Basidia 25-35 X 8-9 µm, tetrasporica, fibulata. Cystidia hymeniales rara, caulocystidia similia. Pileipellis hymeniformis, e cellulis clavatis vel vesiculosis (20-25 X 8-18 µm) , membrana grosse tunicata hyalinaque instructis, pigmento aurantio-brunneo (in KOH) incrustata. Pilocystidia 30-55 X 4- 8 µm, fusiformia vel lecythiformia, membrana grosse tunicata et hyalina instructa. Caulocystidia numerosa, 30-45 X 5-9 µm, pilocystidiis similia. Ad folia putrida Phormii tenacis. Novazelandia. PDD, holotypus.
Gloiocephala tibiicystis belongs in sect. Gloiocephala subsect. Gloiocephala where it is allied with G. mycenoides Singer (1969) and G. epiphylla Massee (1892), The Chilean G. mycenoides differs in forming a densely pubescent, entirely white stipe covered on the base with filiform caulocystidia (and hence the specific epithet), and in forming large conspicuous hymenial cystidia and consistently thin-walled pilocystidia (SGO, holotype!). Gloiocephala epiphylla differs in forming distinctly narrower basidiospores (3.2-4.4 µm diam.), much larger pilocystidia (up to 180 µm long with capitate apex up to 26 µm diam.), and much longer caulocystidia (up to 120 µm long).
New Zealand, North Island, North Auckland, Hokianga Co., Waipoua, 23.vi.l981, Horak (PDD; ZT 1128, isotype).
New Zealand, Canterbury, Ashley, Mt Grey, Kowai River Reserve, 10.xii.1968, Horak (PDD: ZT 68-682).
Pileus to 6 mm diam., hemispheric, soon becoming convex to flat-expanded, centre occasionally depressed, in mature specimens incurved margin dentate-lobate, white, densely covered (hand-lens) with white, strigose hairs. Lamellae none, hymenophore smooth or with few inconspicuous, radially arranged folds, decurrent on stipe, white. Stipe to 4 X 0.3 mm, central, cylindrical, straight to slightly curved, white, pruinose to tomentose overall, fragile, dry, solid, solitary in groups, insititious (byssus or basal disc absent). Odour absent. Basidiospores 9.5-12 X 4- 5 µm, subfusiform to broadly comma-shaped, inequilateral in side profile, smooth, hyaline, inamyloid. Basidia 24-30 X 5-7 µm, constantly bisporic, cylindric to subclavate, sterigmata up to 5 µm long, clamped. Hymenial cystidia absent. Pileipellis a cutis of entangled hyphae (2.5-7 µm) with wart-like, finger-like or more rarely coralloid projections, terminal cells erect, 20-90 µm long, with irregular finger-like projections at base, but smooth towards gradually attenuated tips, non-gelatinous, thick-walled (up to 2 µm diam.), septa clamped. Oleiferous hyphae absent. Stipe tissue monomitic. Stipe cortical hyphae smooth to diverticulate, thick-walled (up to 2 µm), hyaline. Caulocystidia 30-110 X 4-8 µm, scattered, awl-shaped or fusiform with long tapering neck, thick-walled (up to 2 µm).
On rotting leaves of Carex sp. or Uncinia sp. (Cyperaceae). Known from the South Island of New Zealand.
This is the first report of Hemimycena hirsuta (Tode) Singer (sensu Redhead 1981) from New Zealand, which extends its known area of distribution (Eurasia, Morocco in northern Africa, North America) to the southern hemisphere
Pileus to 3 mm diam., to 4 mm high, thimble-like or bell-shaped with a lateral slit, hanging like a street lamp, white, thin-fragile (almost hyaline), smooth. Lamellae none, hymenophore smooth, white. Stipe to 6 X 0.2 mm, central to eccentric, emerging through the lateral slit, cylindrical, whitish to transparent-hyaline overall, very fragile, minutely pruinose, dry, solid, solitary in groups, byssus or basal disc absent. Odour absent. Basidiospores 7-8.5 X 3-3.5 µm, slender elliptic to subcylindric, smooth, hyaline, inamyloid. Basidia 15-20 X 5-6 µm, tetrasporic, cylindric to subclavate, sterigmata up to 4 mm long, clamped. Hymenial cystidia absent. Pileipellis a cutis of cylindric to subfusiform hyphae (3-) 5-16 µm diam., non gelatinous, thin-walled, smooth (except with few, scattered prong-like projections up to 7 µm long), hyaline, inamyloid, clamped. Pilocystidia absent. Oleiferous hyphae absent. Stipe tissue monomitic. Stipe conical hyphae 3-15 µm, cylindric, parallel, non-gelatinous, thick-walled (up to 2 µm), hyaline, inamyloid, clamped. Caulocystidia scattered, 20-40 µm long, awl-shaped or subfusiform with tapering neck, hyaline, walls up to 1 µm thick.
On rhachis of Dicksonia sp. (tree-fern). Known from North Island of New Zealand.
Pileus usque ad 3 mm latus, usque ad 4 mm altus, cylindricus vel campanulatus, albus, fragilis, glaber. Lamellae nullae, hymenium glabrum, album. Stipes usque ad 6 X 0.2 mm, centralis, eccentricus, vel lateraliter affixus, cylindricus, albidus, transparenter hyalinus, fragillimus, minute pruinosus, siccus. Inodorus. Basidiosporae 7-8.5 X 3-3.5 µm, ellipticae vel subcylindricae, leves, hyalinae, inamyloideae. Basidia 15-20 X 5-6 µm, tetrasporica, fibulata. Cystidia hymeniales nulla. Pileipellis cutis ex hyphis cylindraceis vel subfusiformibus cutem formantibus, (3-) 5-16 µm latis, baud gelatinosis, membrana hyalina, tenuitunicata glabraque instructa. Pilocystidia nulla. Caulocystidia rara, 20-40 µm longa, acuto-fusiformia. Ad folia putrida Dicksoniae. Novazelandia. PDD, holotypus.
Hemimycena reducta is characterised by forming very small, bell-shaped basidiomes with a smooth hymenophore and a central to eccentric stipe that emerges through a slit in the pileus. In addition, the pileipellis is formed from nearly smooth hyphae (not a Rameales-structure), hymenial cystidia and pilocystidia are lacking, and caulocystidia are awl-shaped and often setoid. In the field, basidiomes are suggestive of those of Rimbachia Patouillard (1881) and Calyptella Quelet (1886), however, a true stipe is not formed by members of either of these genera. The habit of H. reducta is also suggestive of the genus Cymatella Patouillard (1889), however, in the latter genus basidiomes develop pigmented stipes, and have fusoid basidioles and Rameales-type pileipelli.
Hemimycena reducta differs from other Hemimycena species with reduced hymenophore [such as H. minutissima Desjardin (1991), H. hirsuta (Tode) Singer (1986), Helotium cyphelloides Redhead (1982), Helotium nebulophilum Redhead (1982), Helotium circulare (Singer) Redhead (1984)] by the combination of lacking pilocystidia, lacking numerous diverticula on pileipellis hyphae, and having tetrasporic basidia.
Holotype
New Zealand, North Auckland, Omahuta, 11.v. 1981, Horak (PDD; ZT 543, isotype).
Pileus usque ad 3 mm latus, minutus, convexus subcupuliformis, albus, velutinus vel minute substrigosus, siccus. Lamellae rarae, subvenosae vel nullae, hymenium glabrum, album, velutinosum. Stipes usque ad 10 X ad 0.2 mm, centralis, cylindricus, apicaliter albidus, basin versus niger, ad apicem pruinosus, deorsum levis, siccus, tenax. Inodorus. Basidiosporae 6-7.5 X 2.5-3 µm, subellipticae vel virguliformes. leves, hyalinae, inamyloideae. Basidia 15-20 X 5 µm, tetrasporica, fibulata. Cystidia hymeniales rara, 30-50 X 5-6 µm, fusiformia apice subglobosa (up to 6 µm diam.), tenuitunicata, hyalina. Pileipellis hymeniformis e cellulis clavatis vel subfusiformibus, 15-30 X 8-15 µm, dense diverticulata tenuitunicata, hyalina. Pilocystidia numerosa, 35-45 X 7-12 µm, cystidiis hymeniales similia. Caulocystidia rara, 15-25 X 5-7 µm, cystidiis hymenialibus similia. Ad folia putrida Nothofagi cliffortioides. Novazelandia. PDD, holotypus.
On rotten leaves of Nothofagus solandri var. cliffortioides (Hook. f.) Poole. Known from South Island of New Zealand.
Pileus to 3 mm diam., at first convex, becoming flat-expanded, finally saucer-shaped with upturned margin, white, velutinous to minutely hairy-strigose (under hand-lens), dry, very fragile. Lamellae none, hymenophore smooth or exceptionally with very low venose folds, white, velutinous under hand-lens. Stipe to 10 X to 0.2 mm, central, cylindrical, horsehair-like, apex whitish, lower 3/4 black, apex minutely pruinose, otherwise glabrous, dry, solid, elastic, solitary in groups, insititious (basal disc or byssus absent). Odour absent. Basidiospores 6-7.5 X 2.5-3 µm, slender-elliptic with attenuated base or comma-shaped, smooth, hyaline, inamyloid. Basidia 15-20 X 5 µm, tetrasporic, cylindric to subclavate, sterigmata up to 2 µm long, clamped. Hymenial cystidia scattered, 30-50 X 5-6 µm, fusiform with subglobose apex (up to 6 µm diam.), thin-walled, hyaline. Pileipellis hymeniform, composed of clavate, subfusiform or irregularly shaped broom-cells, 15-30 X 8-15 µm, setulae wart-like or short cylindric (1-4 X 0.5-1 µm), non-gelatinous, thin-walled, hyaline, occasionally with brown intraparietal or encrusting pigment (in KOH), clamped. Pilocystidia numerous, 35-45 X 7-12 µm, apex 5-7 µm diam., shape and structure as hymenial cystidia. Oleiferous hyphae absent. Stipe tissue monomitic. Stipe cortical hyphae cylindric, 3-6 mm diam., parallel, with dark brown intraparietal and/or encrusting pigment, weakly dextrinoid in stipe apex. Caulocystidia scattered, 15-25 X 5-7 µm, shape and structure as hymenial cystidia.
Marasmius meridionalis belongs in sect. Epiphylli subsect. Eufoliatini Singer (1976). It is characterised by forming very small basidiomes with smooth hymenophores, fructifications on leaves of Nothofagus, small basidiospores, weakly dextrinoid tissue in the stipe apex, and a pileipellis composed of setulose cells plus capitate pilocystidia. Marasmius meridionalis differs from all other taxa in the section in the combination of a smooth hymenophore and setulose pileipellis cells. The European M. epiphylloides (Rea) Saccardo & Sydow is similar in some respects, but differs in forming well-developed lamellae, fructifications on leaves of Hedera, and in having thick-walled pilocystidia, inamyloid stipe tissues, and basidiospores 11-15 µm long
Holotype: New Zealand, Canterbury, Ashley, Mt Grey, Kowai River Reserve, 10.xii.l968, Horak (PDD; ZT 68-681, isotype).
New Zealand, North Island: North Auckland, Hokianga Co., Waipoua Kauri Reserve, 12.v.1983, R. H. Petersen 55634 (TENN 44402, holotype). Gisborne, Urewera N. P., near Headquarters, 31.v. 1981, Horak (PDD; ZT 849).
Description of Second New Zealand Record
Pileus to 3 mm, convex to flat-expanded, old specimens also with depressed centre, subumbilicate, whitish, minutely pruinose, margin subciliate. Lamellae none, hymenophore smooth or occasionally with a few low radially arranged folds, whitish. Stipe to 4 X to 0.2 mm, central (rarely eccentric), cylindrical, straight or curved, whitish to hyaline, minutely pruinose overall, dry, solid, solitary in groups, insititious (byssus or basal disc absent). Odour absent. Basidiospores 12-16 X 4.5-5.5 µm, subfusiform (reminiscent of boletoid spores), inequilateral in profile, often with significant supra-apicular depression, smooth, hyaline, inamyloid. Basidia 23-30 X 8-11 µm, tetrasporic, subclavate, clamped. Hymenial cystidia inconspicuous, lageniform. Pileipellis hymeniform, composed of subcylindric, clavate or broadly fusiform ceils, 30-60 X 8-20 µm, non-gelatinous walls up to 1µm diam., hyaline, smooth. Pilocystidia scattered, up to 90 X up to l6 µm, fusiform with tapering, rounded neck, thin-walled, hyaline. Margin of cap with scattered, bristle-like, cylindrical 'hairs', 3-7 µm diam., subgelatinous, conpicuously thick-walled (up to 3 µm diam.), with distinctive lumen at apex, often encrusted with colourless matter. Oleiferous hyphae absent. Stipe tissue monomitic. Stipe conical hyphae 3-8 µm diam., parallel, cylindric, hyaline, smooth. Caulocystidia numerous, 15-50 X 8-18 µm, polymorphic, ranging in shape from cylindric to broadly fusoid, walls up to 1.5 µm thick, hyaline, clamped.
On rotting leaves of Astelia sp. (Asteliaceae), already infecting healthy leaves and consistently associated with leaf spots. Known from North Island of New Zealand.
This represents a second New Zealand record for this tiny species reported first from Waipoua Kauri Reserve, Hokianga Co. (Desjardin and Petersen 1989).
The material described above differs slightly from the holotype in showing larger pilei, longer pilocystidia, and the presence of scattered bristle-like marginal hairs. Gloiocephala Massee, Grevillea 21:33(1892)

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18 March 2001
5 March 2010
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