Zotov, V. D. 1971: Zoysia Willd. (Gramineae) in New Zealand. New Zealand Journal of Botany 9: 639–644.
Details
Zotov, V. D. 1971: Zoysia Willd. (Gramineae) in New Zealand. New Zealand Journal of Botany 9: 639–644.
Article
Taxonomic concepts
Descriptions
Plant perennial. Rhizome internodes mostly in series of three — 2 very short and 1 long — the three internodes 1-2 cm long. Cataphylls rounded to more or less truncate, Ungulate, distinctly umbonate. Erect culms arise from short internodes, 2-5 cm high above ground, branching several times at and above ground. Branching intravaginal; 2-3 internodes elongated, uppermost, 0.7-3 cm long. Leaf sheath 5-15 mm long, glabrous finely striated. Ligule 0.2-0.4 mm long, ciliar. Leaf collar distinct, more or less long-ciliate, hairs up to 5 mm long, cushion based. Blade more or less distinctly divergent, folded, subulate, glabrous. Vascular bundles about 11. Sclerenchyma bilateral, interrupted between bundles; connected with primary bundles, more or less free from secondary bundles. Peduncle 1-2 cm (seldom more) long, finely striated and minutely scabrid above. Spikelet solitary (occasionally 2) terminal, erect, 3.5-5.0 mm long, lanceolate. Glume 4 mm long, ovate indurated, minutely hairy above, more or less golden to golden-brown, about 7- nerved, nerves very weak. Lemma about 3.5 mm long ovate-lanceolate, more or less conduplicate, 1-nerved, firm hyaline, glabrous. Anthers about 2 mm long. Stigmas long, plumose (Fig. 1).
Plant perennial with extensive wiry rhizomes. Rhizomes mostly in series of three internodes — 2 very short and 1 long — the three internodes 1-3 cm long. The longer internode subtended by longer cataphyll. Cataphylls rounded to truncate, lingulate, more or less distinctly umbonate. Erect culms arise from short internodes, 4-10 cm high above ground, branching several times at and above ground. Branching intravaginal; 2-3 internodes elongated, uppermost mostly 2-5 cm long. Leaf sheath 1-3 cm long, glabrous, finely striated. Ligule 0.2-0.5 mm long, ciliar. Leaf collar distinct, more or less ciliate with long soft cushionbased hairs. Blade up to 7 cm long, more or less flat 1-1.5 mm wide, tapering, more or less finely striate on both sides, glabrous. Vascular bundles mostly about 15. Sclerenchyma bilateral, interrupted between vascular bundles, connected with primary bundles, more or less free from secondary bundles. Peduncle 2-5 cm long, glabrous. Racemes 7-12 mm long of 3-5 spikelets. Rachis and pedicels minutely scabrid. Pedicels 0.5-1 mm long, spikelets about 4 mm long, lanceolate. Glume 4 mm long, ± ovate, indurated with thin margins minutely hairy above and on margins, more or less golden ochreous, about 7-nerved, nerves weak, awn 0-5 mm long. Lemma about 3.5 mm long ovate-lanceolate, glabrous. Anthers about 2 mm long. Stigmas long, plumose (Fig. 1).
Plant perennial. Rhizome internodes mostly in series of three — 2 very short and 1 long — the three internodes 1-2 cm long. Cataphylls truncate, lingulate, almost quite bladeless. Erect culms arise from one or both short internodes, 8-15 cm high above ground, branching several times at and above ground. Branching intravaginal; 2-3 internodes elongated, uppermost 4-7 cm long. Leaf sheath 1-3 cm long, glabrous finely striated. Ligule a line of hairs 0.3-0.5 mm long. Leaf collar distinct, more or less densely long-ciliate, hairs soft, cushion-based. Blade more or less distinctly divergent rolled in bud, flat at maturity, up to 5 cm long 2-3 mm wide, tapering, finely striate on both sides, glabrous. Vascular bundles about 25. Sclerenchyma bilateral, interrupted between vascular bundles, connected with primary bundles, more or less free from secondary bundles. Peduncle 4-7 cm long, glabrous. Racemes 8-16 mm long, of 6-12 appressed spikelets. Rachis and pedicels minutely scabrid. Pedicels 0.5-1 mm long. Spikelets about 4 mm long, lanceolate. Glume 4 mm long, ovate, indurated with thin margins, minutely hairy above and on margins, stramineous, about 7-nerved, nerves very weak. Lemma about 3.5 mm long, ovate-lanceolate, more or less conduplicate, 1-nerved, firm, hyaline, glabrous. Anthers about 2 mm long. Stigmas long, plumose (Fig. 1).
Planta perennis. Rhizomatis internodia persaepe in tripartita serie disposita — duo brevissirna, unum longum — haec tria internodia 1 - 2 cm longa. Internodium longius longiore cataphyllo est subtentum. Cataphylla truncata, lingulata, paene omnino laminis carentia. Culmi erecti, 8 - 1 5 cm alti, oriuntur ex axillis brevium cataphyllorum vel unius vel amborum, aliquoties ramos emittentes ad soli superficiem vel supra. Ramificatio intravaginalis; 2 - 3 internodia elongata, supremum 4 - 7 cm longum. Folii vagina 1 - 3 cm longa, glabra, subtiliter striata. Ligula ciliaris, 0.3 - 0.5 mm longa. Folii collum distinctum plus minusve dense longeciliatum, pilis mollibus basi pulviniformibus. Lamina plus minusve distincte divergens, in gemma involuta, ad maturitatem plana, ad 5 cm longa, 2 - 3 mm lata angustata, in utraque facie subtiliter striata, glabra. Fasciculi vasculares circa 25. Sclerenchyma bilaterale, inter fasciculos vasculares interruptum conjunctum cum fasciculis primariis, ab fasciculis secundariis plus minusve disjunctum. Pedunculus 3 - 7 cm longus, glaber. Racemi 8 - 1 6 mm longi ex 6 - 12 spiculis appressis constantes. Rhachis pedicellique minute scabridi. Pedicelli 0.5 - 1 mm longi. Spiculae circa 4 cm longae, lanceolatae. Gluma 4 mm longa, oyata, indurata, marginibus tenuibus, minute pilosa supra et in marginibus, straminea, circa 7-nervis; nervi obscurissimi. Lemma circa 3 - 5 mm longum, ovato-Ianceolatum, plus minusve conduplicatum, 1-nerve, firmum, hyalinum, glabrum. Antherae ciica 2 mm longae. Stigmata longa, plumosa.
Plant perennial, with extensive monopodial rhizomes. Culms few, noded. Leaf sheath non-auriculate; ligule ciliar; collar distinct, more or less long ciliate with cushion based hairs. Leaf blade linear to linearlanceolate, tapering. Silica cells round or slightly concave on upper and lower edges. Long cells finely papillate. Microhairs broad clavate; terminal cell more or less hemispherical; pediment somewhat bulbous. Vascular bundles of first order spaced by several bundles of second order. Chlorenchyma concentric. Inflorescence a raceme. Spikelets 1- flowered, disarticulating entire above pedicels. Lower glume absent. Upper glume laterally compressed, infolding the floret, indurated, more or less mucronate. Lemma entire, firm membranous, hyaline, 1-nerved. Palea Ungulate, nerveless. Lodicules absent. Stamens 3, ovary glabrous. Styles 2, distinct, approximate elongate. Stigmas long, emerge apically. Fruit elliptic, laterally compressed, free. Seed free within pericarp. Starch grains simple. Embryo about half as long as seed. Scutellum cochleariform. Epiblast broad, conspicuous. Plumule distinctly stalked. Coleorhim more or less free of scutellum. Chromosomes small (2-3 mcm long); basic number 10.
Identification keys
Racemes with 15 - 30 spikelets; culms 12 - 25 cm tall; lea blades flat (not in New Zealand)
2
Racemes reduced to 1 (rarely 2) spikelets; culms 2 - 6 cm tall; leaf filiform, terete
3
Racemes with 3 - 5 spikelets; culms 5 - 10 cm tall; leaf blades filiform ± folded to flat 0.8 - 1.5 mm wide
Reproduced with permission from New Zealand Journal of Botany and The Royal Society of New Zealand Te Apārangi.
Cited scientific names
Metadata
0dbcdc36-fd1f-4539-9252-1d5f413d6413
reference
Names_Plants
19 July 2005