Thallus heteromerous, foliose, lobate, dorsiventral, lobes 0.5-5(-8) cm diam. Upper surface corticate, cortex paraplectenchymatous, smooth or ± wrinkled or plicate, glabrous, matt or shining or ± scabrid or tomentose or pruinose, bluish-, brownish or yellowish-green or reddish-brown, terricolous or muscicolous or corticolous. Medulla white, loosely interwoven. Photobiont blue-green, , or green, . Lower surface decorticate, arachnoid, ± whitish or brownish, ± distinctly veined. Veins flat or raised, rhizinate. Rhizines short or long, pale or dark, simple to fasciculate, often with an anchoring squarrose tuft. Apothecia round or irregular, marginal, horizontal or on vertical, ascending lobules, often with margins reflexed, exciple areolate-tomentose, or ± lacking, disc red-brown to brown-black, epruinose, margins flesh-coloured to pale brownish, often ± verrucose. Epithecium red-brown. Hymenium colourless to pale brownish. Hypothecium colourless to brown. Asci clavate-cylindrical, 8-spored with a well-defined apical ring structure. Ascospores fusiform or acicular, 3-9-septate, colourless or brown.