[GS] Pileus 2.5-3.5 cm. diam., reddish brown, orbicular to ovate with downrolled margin, subfibrillose; flesh concolorous, tough gelatinous. Gills decurrent, moderately crowded, shallow, fawn. Stipe 0.5-1 X 0.2-0.7 cm., lateral or markedly excentric, concolorous, velutinate. Spores 7-8um diam., non-amyloid, globose, thin-walled (Fig. 1/47, p. 10). Hymenophoral trama and cuticle of loosely woven, partly gelatinized hyphae. HABITAT: on fallen JVothofagus menziesii, Nelson, 6.7.1949, A. Crawford in Stevenson (type). [Buchanan et al] In his revision of the Agaricales of New Zealand, Horak (1971) appears to have been confused by the same epithet having been applied by Stevenson (1964) to two pleurotoid fungi, namely Lentinellus crawfordii and Resupinatus crawfordii. In his list, Horak (1971) cited Resupinatus (Lentinellus) crawfordii, seeming to indicate that he considered the two species to be the same. He noted that 'the identification of Stevenson cannot be confirmed because carpophores of the type material are sterile.' As shown above, Lentinellus crawfordii is easily recognised and a good species. However, the type material of R. crawfordii (Stevenson 718, on fallen Nothofagus menziesii, Nelson, A.Crawford, 6. vii. 1949, K) is certainly in a poor state, and little detail of the hymenium could be gleaned. Spores were found, however, which were short cylindric to slightly allantoid and strongly amyloid, and there were densely staining amyloid remains of spores clinging to the hymenium. The cheilocystidia and pleurocystidia are difficult to determine accurately, but the lamellar edge is sterile and appears to consist of short, clavate cheilocystidia. The trama is dense, of more or less parallel, thickish-walled hyphae of 3-5 um diameter. The subhymenium is cellular, very densely compacted of narrow, somewhat thick-walled hyphae. The context is of very narrow (2-3 um in diameter), loosely interwoven hyphae with large clamp connections. The pileipellis appears to be only little differentiated, with repent hyphae with coiled, slightly inflated hyphal endings. The stipe is short and stout, with distinctly velvety to fibrillose surface. The fibrils are made up of bundles of narrow hyphae, or the surface of the stipe may be covered with short caulocystidia. The fungus can be readily identified as a species of Panellus. The size of the spores, the absence of distinctive cheilocystidia, and the thin-walled pileipellis as well as the more robust form of the basidiome distinguish it from Panellus stypticus (Bull.: Fr.) P. Karst. which is also present in New Zealand and described by Petersen and Bermudes (1992). The new combination proposed for this fungus is Panellus crawfordii. [JAC] The pileipellis is a dense cutis of long thin clamped hyphae over a gelatinized subpellis. Spores amyloid length=6.0–9.4µm (µ=7.4, σ=0.72), width=3.3–4.5µm (µ=3.9, σ=0.31), Q=1.6–2.2µm (µ=1.90, σ=0.16), n=20. Lamella without cystidia, trama with long elements like pileipellis. The macro and micro moprhology confirm my earlier suspicion that this is yeat another synonym of Scytinotus longuinquus.

The identification of Stevenson cannot be confirmed because carpophores of the type material are sterile.

[Notes from Kew Type specimen, PRJ 2010] Kew images.

Pileus 2.5-3.5 µm diam., reddish brown, orbicular to ovate with downrolled margin, subfibrillose; flesh concolorous, tough gelatinous. Gills decurrent, moderately crowded, shallow, fawn. Stipe 0.5-1 x 0.2-0.7 cm, lateral or markedly excentric, concolorous, velutinate. Spores 7-8 µm diam., non-amyloid, globose, thin-walled (Fig. 47). Hymenophoral trama, 1 µm cuticle of loosely woven, partly gelatinized hyphae. Spors smooth.

On fallen Nothofagus menziesii. Nelson, 6.7.1949, A. Crawford in Stevenson (type).

Pileus 2.5-3.5 µm diam., rubido-brunneus, orbicularis usque ovatus, margine deorsum involuto, subfibrillosus; caro concolor, tenaciter gelatinosa. Lamellae decurrentes, modice confertae, haud altae, hinnuleae. Stipes 0.5-1 x 0.2-0.7 cm, lateralis vel manifeste excentricus, concolor, velutinosus. Sporae 7-8 µm diam., haud amyloideae, globosae, parietibus tenuibus. Trama hymenophoralis et cuticula ex hyphis laxe intertextis ex parte gelatinatis sistentes.

Typus: A. Crawford in Stevenson 718.