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Marasmiellus omphaloides G. Stev. 1964

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This is indigenous
Threat status: Data deficient
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Marasmiellus omphaloides G. Stev., Kew Bull. 19 33 (1964)
Marasmiellus omphaloides G. Stev. 1964

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Endemic
Present
New Zealand
Political Region

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G. Stev.
G. Stev.
1964
33
ICN
Marasmiellus omphaloides G. Stev. 1964
NZ holotype
species
Marasmiellus omphaloides
on fallen wood, Waikaneae, 1.1.1951, Stevenson (type),

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omphaloides

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Marasmiellus omphaloides G. Stev. 1964

[GS] Pileus 11-17 mm. diam., dull white, thin, dry, hemispherical at first becoming plano-concave, somewhat fluted at margin. Gills adnate-decurrent, creamy white, distant, occasionally forked and with folds between. Stipe 2 cm. x 1 mm., tapering downwards, cream immediately under the gills, fawn to purple brown below, solid, tough. Spores 11-12 X 4-4.5um, non-amyloid, thin-walled. Hymenophoral trama non-amyloid. Cuticle of woven repent non-amyloid hyphae with clamp connections, with sparse superficial dermatocystidia which are hair-like, amyloid and thick-walled. (Fig. 2/14, P- 37) [EH] This species belongs to a group of apparently related Marasmiellus like fungi known from New Zealand which possess subamygdaliform, smooth spores and long, thick-walled, yellow-brown, hair-like dermatocystidia in the cuticle. To our knowledge there is no existing genus where these fungi could be adequately accommodated.[JAC] non Marasmiellus omphalodes (Berk & Curt.) Singer (twice). This material attacked by an Aspergillus. The gills have dried yellow/orange like some Campanellus. No sign of sterile gill edge. The cap appears minutely felty. Stipe with indistinct macro-visible cystidia. Stem arising from a superficial slightly enlarged base on wood with fine indistinct brown bristles. Pilleipellis with isolated clusters of yellow, thick-walled, fragile/fracturing hairs, associated with shorter, nearly vesiculose, and clamped elements (hydropoid?). Stipe with similar hyaline hairs (and numerous allantoid spores of a tremellaceous fungus). Spores somewhat collapsed. No cheilocystidia. No tissue amyloid or dextrinoid. Material infested with globose echinulate spores of the Aspergillus. Stipitipellis thick-walled, with swollen septa, clamps not observed. 4-spored basidia observed (mostly collapsed) but some evidence of 2-spored given range in spore size. No clamps observed in pileipellis or hymenium but tissue highly disorganised. Spores length=10.8–14.6µm (µ=12.8, σ=0.96), width=4.5–6.2µm (µ=5.4, σ=0.40), Q=2.0–2.9µm (µ=2.38, σ=0.20), n=25. Note that M. candidus ss NZ (marasmiaceae) has hair-like pileocystdia, but not yellow and not thick walled. M. Mt Fyffe (porotheliaceae) has thick walled pileocystidia but they are hyaline and present also as cheilocystidia. Previous idents of M. omphalodes (PDD 95810, PDD 87725, lyophyllaceae ) are now questionable because they do not have yellow pilocystidia and posess cheilocystidia. If hydropid then closer to M. Mt fyffe, but that with clear cheilocystidia.

Marasmiellus omphaloides G. Stev. 1964

This species belongs to a group of apparently related Marasmiellus-like fungi known from New Zealand which possess subamygdaliform, smooth spores and long, thick-walled, yellow-brown, hair-like dermatocystidia in the cuticle. To our knowledge there is no existing genus where these fungi could be adequately accommodated.

Marasmiellus omphaloides G. Stev. 1964

[Notes from Kew Type specimen, PRJ 2010] Kew images.

Marasmiellus omphaloides G. Stev. 1964

Pileus 11-17 mm diam., dull white, thin, dry, hemispherical at first becoming plano-concave, somewhat fluted at margin. Gills adnate-decurrent, creamy white, distant, occasionally forked and with folds between. Stipe 2 cm x 1 mm, tapering downwards, cream immediately under the gills, fawn to purple brown below, solid, tough. Spores 11-12 x 4-5.5 µm, non-amyloid, thin-walled. Hymenophoral trama non-amyloid. Cuticle of woven repent non-amyloid hyphae with clamp connections, with sparse superficial dermatocystidia which are hair-like, amyloid and thick-walled. (Fig.14).
On fallen wood, Waikaneae, 1.1.1951, Stevenson (type).
Pileus 11-17 mm diam., sordide albus, tenuis, siccus, primo hemisphaericus deinde plano-concavus, margine nonnihil striato. Lamellae adnato-decurrentes, cremeo-albae, distantes, interdum furcatae, plicis interpositis. Stipes 2 cm x 1 mm, deorsum attenuatus, sub ipsas lamellas cremeus, inferne hinnuleus usque purpureo-brunneus, solidus, tenax. Sporae 11-12 x 4-4.5 µm, haud amyloideae, parietibus tenuibus. Trama hymenophoralis haud amyloidea. Cuticula ex hyphis intertextis repentibus haud amyloideis fibulatis sisteus, dermatocystidiis superficialibus sparsis filiformibus amyloideis parietibus crassis praedita.
Typus: Stevenson 801.

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Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev.
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. (1964)
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. (1964)
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. (1964)
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. (1964)
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. (1964)
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. (1964)
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. 1964
Marasmiellus omphaloides G. Stev. (1964)

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Marasmiellus omphaloides G. Stev. 1964
New Zealand
Kaikoura
Marasmiellus omphaloides G. Stev. 1964
New Zealand
Mid Canterbury
Marasmiellus omphaloides G. Stev. 1964
New Zealand
Nelson
Marasmiellus omphaloides G. Stev. 1964
New Zealand
North Canterbury

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taxonomic status
This Marasmiellus-like fungus could not be adequately accommodated in any existing genus, fide Horak 1971b, p. 442. Sequence evidence (ITS for PDD95810) suggests this is related to Marasmiellus ramealis, distant from core marasmiellus and perhaps better referred to Collybiopsis.
typification
on fallen wood, Waikaneae, 1.1.1951, Stevenson (type),

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1cb19232-36b9-11d5-9548-00d0592d548c
scientific name
Names_Fungi
1 January 2000
15 December 2003
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