As for but differing in the following respects: Thallus without isidia or soredia, containing only zeorin and pannarin (KC-, Pd+ orange). Apothecial margins brown to black. Hymenium 120-250 µm tall. Ascospores 3-septate or muriform with thick, transverse septa.

Thallus crustose, heteromerous, yellowish-, greenish- or whitish-grey, thin or rather thick c. 50-150 µm, surface smooth, wrinkled, verrucose-papillate or with isidia or soredia. Photobiont present in upper 60 µm of thallus, green, subspherical, 6-12 µm diam., sp. Apothecia biatorine, sessile to subpedicellate, 0.5-5.0 mm diam., disc plane to convex, brown to black, sometimes with a greyish, beige or whitish pruina, margins yellowish, brown or black, usually paler than disc, sometimes darker, disc and margins usually slightly glossy. Epithecium c. 15 µm thick, usually diffusely orange-brown, sometimes with pigments or granular inclusions. Hymenium 80-260 µm tall, colourless, inspersed with oil droplets, c. 1-7 µm diam., I reaction variable. Paraphyses 1 µm thick, indistinctly septate, parallel, little branched, in epithecium more branched and anastomosing and slightly thickened (2-3 µm). Hypothecium usually 30-50 µm thick, usually less inspersed than hymenium. Ascospores 1-8 per ascus, colourless, 1-septate, to multiseptate, with thin septa, ellipsoid, ovoid or ± subspherical, straight or curved, sometimes with thickened apices, and a prominent, smooth or warted epispore. Chemistry: Zeorin and usnic acid or pannarin.