Annual or perennial herbs, sometimes somewhat woody. Lvs usually opposite, rarely alternate, simple and entire to dentate or lobed, or 1-3-ternate or pinnate. Capitula pedunculate, solitary or in loose cymes. Involucral bracts in 2-(4) rows; outer row usually foliaceous; inner rows membranous. Receptacle flat or slightly convex; scales present. Outer florets ♀ or sterile, ligulate, usually yellow, sometimes mauve, rose, pink, white or marked with red toward base. Inner florets ☿, tubular. Achenes all similar, orbicular, oblong to linear, compressed; lateral angles usually with entire or dentate wings, sometimes not winged; pappus 0, reduced to a small corona, or of 2 antrorsely barbed or smooth awns.