Annual, biennial or perennial herbs; hairs various. Basal lvs often long-petiolate; cauline lvs short-petiolate or sessile, often amplexicaul. Infl. usually paniculate, composed of ebracteate cymes. Calyx deeply lobed, often to the base, persistent and often enlarged at fruiting. Corolla tube short, cylindric or funnelform; lobes ± patent, broad; scales 5, closing throat. Stamens included, inserted at or above middle of tube. Style included, < calyx. Nutlets ovoid to subglobose, glochidiate, sometimes with distinct margin or disc, slightly concave to convex on outer surface.