Biennial to perennial taprooted herbs. Hairs simple. Stems erect, leafy. Lvs simple, cordate, crenate, serrate or dentate. Racemes ebracteate. Sepals erect, the inner deeply saccate. Petals purple, rarely white. Stamens 6, the inner 4 with broad filaments. Lateral nectaries 4, paired beside bases of lateral stamens; median nectaries 0. Style short; stigma 2-lobed. Silicle lanceolate or elliptic, large, flat, latiseptate, dehiscent, raised above the sepal scars on a short or long carpophore; valves thin with fine net venation; septum silvery. Seeds large, circular, flattened, winged, in 2 rows per locule.