Primary squamules persistent or disappearing, medium 1-4 × 0.5 mm, digitate-laciniate, crenate, ascending, flat or involute, scattered or in tufts, rarely forming cushions, upper surface glaucescent to olive-brownish, rarely whitish-glaucescent, lower surface white or base browning or red-brown, esorediate. Podetia slender, growing from upper surface of primary squamules, base persistent or dying away and growth continuing apically, 10-110 mm tall, to 5 mm diam., cupless or bearing cups, cups perforate, open to interior of podetia, flaring abruptly, when cupless, branches radiately or sympodially developed, margins of cups becoming repeatedly proliferate, axils of branches commonly perforate, tips of proliferations with cups, blunt, pointed or with apothecia, esorediate. Cortex continuous or suncontinuous, smooth or finely areolate with a white reticulum, shining or dull, opaque, glaucescent or pale or olive-green or brownish to red-brown, dying parts black. Apothecia small, at tips of podetia, or on short corymbose branchlets, brown or red-brown. Chemistry: Cortex K-, KC-, Pd-. Squamatic (UV+) and ± barbatic acids.