Thallus heteromerous, granular, microphylline or squamulose to subfoliose, often erect, upper cortex well-developed, ± loosely attached to substrate by medullary hyphae or by a prothallus, isidia present or absent, margins of thallus ± pale, corticolous. Photobiont green, . Apothecia sessile, biatorine, ± convex, brown or reddish-brown with a thin, pale proper margin, disappearing with age. Paraphyses distinct, simple, apices not noticeably thickened, conglutinate. Asci clavate, 8-spored. Ascospores simple, colourless, long-ellipsoid. Hypothecium massive, 150-250 µm thick, intensely gelatinised, appearing chondroid.