Perennial, usually hispid herbs. Basal lvs with petiole long and welldefined; cauline Ivs sometimes sessile. Infl. of short, ebracteate terminal cymes. Calyx 5-lobed, often divided nearly to base, accrescent. Corolla tube cylindric in lower 1/2, tubular-campanulate or funnelform above; limb of 5 short lobes; scales present, usually with papillate margins. Stamens included, inserted around middle of tube. Style exserted; stigma very small. Nutlets ovoid, erect or curved, usually rugose or verruculose, occasionally smooth; base concave with thickened collar.