Primary squamules larger than in , 5-10 mm long and 2.5-5 mm wide, incised-lobate, persistent or disappearing, upper surface yellowish-glaucescent, lower surface white, pale to dark brown or blackened centrally, occasionally sorediate. Podetia 20-85 mm tall, 2-10 mm wide, terete, with or without cups, cups gradually flaring from base, more irregular than in , marginal proliferations prominent, sterile podetia tapering towards apices, sides often fissured and split. Cortex densely farinose-sorediate, yellowish-grey or yellowish, often ± corticate or squamulose at base. Apothecia red, sessile, on cup margins. Chemistry: Cortex K-, C-, KC+ yellowish, Pd-. Squamatic (UV+) and usnic acids.