Unarmed, annual or perennial herbs, or low subshrubs. Lvs alternate, simple, rather large, membranous, often hairy, at least when young, entire, toothed or lobed, petiolate. Fls tending to form leafy cymose infls, ± erect. Calyx 5-toothed but teeth often small, circumscissile. Corolla large, usually narrow- to broad-funnelform, plicate, white to purple, 5-toothed. Stamens 5, inserted near base of corolla tube, included to slightly exserted. Stigma 2-lobed. Capsule dehiscing by 4 valves, ± spinose, erect or pendent. Seeds many, of moderate to fairly large size, to 6 × 4.5 mm.