Thallus foliose, lobate, dorsiventral, heteromerous, loosely to moderately firmly attached to substrate, corticolous. Lobes 0.4-11.0 mm broad, short and rounded to elongate, ± flat. Upper surface brown to brownish-black, with or without isidia, pseudocyphellae or soredia. Lower surface rhizinate. Rhizines mostly unbranched, rarely branched at tip. Apothecia rare in species producing asexual propagules, lecanorine, sessile to subpedicellate, disc smooth, imperforate, margins entire, flexuose at first, entire to variously notched and incised or crenulate with age, concolorous with thallus. Cortex lacking any reaction with HNO3.