Notes: Skierka species are tropical and autoecious (Mains 1939c, Cummins & Hiratsuka 2003). All sori are subepidermal and deep-seated; non-catenulate teliospores are extruded in hair-like columns. Skierka has long held an isolated placement laced Skierka in a separate subfamily or tribe, respectively, in the Pucciniaceae; Cummins & Hiratsuka (2003) treat it as incertae sedis within the rusts. Mains (1939c) hypothesised that Skierka represented an intermediate taxon between the Melampsoraceae and Pucciniaceae (equivalent to the subordinal ranks Melampsorineae and Raveneliineae/Uredinineae, under the present classification), a position largely congruent with our placement.