Evergreen woody liane, usually climbing Over shrubs or small trees; dioecious. Stems up to 5 m long, 5-10 mm diam., ribbed and sparsely hairy when young, glabrous when mature, ascending or spreading. Leaves 3- foliolate, thin to coriaceous, green to grey-green. Leaflets 14-55 × 9-28 mm, simple to pinnate; simple leaflets lanceolate, broadly ovate or deltoid, entire, crenate, serrate or deeply pinnatifid to bipinnatisect; pinnae linear-oblong to broadly ovate; apices of leaflets, pinnae, crenations and serrations apiculate; bases attenuate to truncate, sometimes slightly cordate; petioles and petiolules 10-60 mm long, channelled, glabrous to sparsely hairy, twin- ing. Inflorescences axillary; flowers solitary, or 2- 10 clustered in leaf axils, or in dichasia1 cymes of 5-10; flowers unisexual, 10-45 mm diam. Pedicels 25-95 mm long, moderately to densely hairy; bracts 3-13(-18) × 14(-8) mm, inserted about middle of pedicel, usually linear to elliptic, occasionally broadly elliptic, connate, abaxial and adaxial surfaces ± glabrous to moderately hairy, apex acute to obtuse. Male flowers: sepals 5-6, 10.0-25.0 × 3.0- 13.0 mm, narrowly to broadly elliptic, lanceolate or ovate, cream, often flushed red-brown at base, abaxial and adaxial surfaces with moderate to dense pilose or villous hairs, rarely glabrous on the adaxial surface, apex subacute to obtuse; stamens 20-37; filaments 3-1 1 mm long, c. 0.9 mm wide, glabrous, linear, often tapering noticeably toward apex; anthers 2.4-3.5 mm long, 0.4-0.5 mm wide, linear-oblong, cream; receptacle 0.6-2.2 mm high, 1.4-2.2 mm wide, ± glabrous to moderately hairy; carpelodes absent. Female flowers: sepals 54(-7), 13.0-25.0 × 3.8-7.0 mm, narrowly elliptic to broadly elliptic, lanceolate to ovate, cream, often flushed red at base, abaxial and adaxial surfaces with moderate to dense pilose or villous hairs, apex subacute to obtuse; car- pels 35-48; ovary 0.5 × 0.2 mm, densely hairy; style 4-5 mm long; pappus hairs 5-6 mm long, dense, white, ascending; stigma 0.4-2.5 mm long, curved, papillate on adaxial surface, abaxial surface smooth; receptacle 1.8-2.3 mm high, 1.2-1.5 mm wide, moderately hairy; staminodes 1-6, in outer whorl; filaments 3.4-5.2 mm long: anthers c. I mm long, malformed, barren. Achenes 3.0-4.0 mm long, 1.4- 1.5 mm wide, 0.6-0.8 mm thick, sparsely to moderately hairy, dark brown to light brown, rarely red-brown; style 8-20 mm long, Pappus hairs spreading. FL SepMar; FT Oct-Apr.

Evergreen woody liane, usually climbing over shrubs and small trees; dioecious. Stems up to 2 m long, 5-6 mm diam., ribbed and sparsely hairy when young, glabrous when mature, ascending or spreading. Leaves 3- foliolate, subcoriaceous to coriaceous, green. Leaflets 10-30 × 4- 18 mm. simple to pinnate; simple leaflets broadly ovate to ovate, entire, crenate to deeply pinnatifid; pinnae linear-oblong to broadly ovate; apices of leaflets and pinnae apiculate, bases sometimes oblique, attenuate to truncate; petioles and petiolules 10-25 mm long. channelled, glabrous to sparsely hairy, twining. Inflorescences axillary; flowers solitary, or 2-6 clustered in leaf axils, or in dichasia1 cymes of 5-10; flowers unisexual, 12- 35 mm diam. Pedicels 10-40 mm long, moderately to densely hairy; bracts 6.0- 14.0 × 1.7-6.0 mm, inserted about middle of pedicel, often foliaceous, connate, spathulate, often tinged red at base, abaxial surface sparsely to moderately hairy, adaxial surface ± glabrous to sparsely hairy, apex obtuse to rounded. Male tlowers: sepals 6-8, 5.5-20.0 × 1.7-9.0 mm, narrowly elliptic, elliptic, lanceolate to ovate, yellow-green, abaxial surface with pilose to villous hairs, adaxial surface glabrous or very rarely with a few scattered hairs, apex subacute to obtuse; stamens 14-24; filaments 1.8-7.4 mm long, c. 0.75 mm wide, glabrous, linear, often gradually tapering toward distal end or slightly at both proximal and distal ends; anthers 2.0-2.3 mm long, c. 0.6 mm wide, linear- oblong, cream; receptacle 0.5-0.6 mm high, 1.0- 1.1 mm wide, ± glabrous to moderately hairy; carpelodes absent. Female flowers: sepals 6-8.6.0- 15.0 × 3.5-5.0 mm, narrowly elliptic, elliptic, lanceolate to ovate, yellow-green, abaxial surface with pilose to villous hairs, adaxial surface glabrous, apex subacute to obtuse; carpels 30-39; ovary c. 1.2 × c. 0.4 mm, glabrous; style 4.4-5.3 mm long; pappus hairs 5 4 mm long, dense, white, ascending; stigma 0.4-2.7 mm long, curved, papillate on adaxial surface, abaxial surface smooth; receptacle 2.0- 2.2 mm high, 1.0- 1.1 mm wide, moderately hairy; staminodes 6-12, in outer whorl; filaments 4.0- 4.2 mm long; anthers c. 1.0 mm long, malformed, barren. Achenes 3.5-4.4 mm long, 1.5-2.3 mm wide, 1.0- 1.2 mm thick, glabrous, chesmut-brown; style 5-17.5 mm long, pappus hairs spreading, base of style usually glabrous. FL Oct-Jan; FT Dec-Mar.