Colonies amphigenous, mainly hypophyllous, occasionally on petioles, effused and often ascending leaf hairs. Mycelium superficial, pale, branched and anastomosing, smooth or verruculose, septate,1.5-2.5 µm thick; hyphae below conidiophores thicker, 2.5-6 µm and brown, often forming dark brown multicellular plates or stromata on the leaf hairs. Conidiophores arising terminally and laterally on the hyphae, macronematous, single, composed of a mononematous stipe and a complex head of branches bearing conidia. Stipes erect, straight or flexuous, smooth or verruculose especially at base, septate, dark brown, paler towards the apex, up to 425 µm long x 8-11 µm thick at base tapering to 5-6 µm thick at the apex. Primary branches sometimes bearing branchlets, brown, flexuous, up to 75 µm long x 3.5-5.5 ,um thick. Conidiogenous cells polyblastic, integrated, terminal becoming intercalary, sympodial, conidial scars prominent, 1-1.75 µm wide. Conidia solitary, acropleurogenous, pale brown, smooth, 3-4 (-6) septate, usually slightly onstricted at septa, straight or slightly curved, obclavate-cylindric, apex rounded, base obconically truncate, (23-) 25-40 (-51) x 4.5-5 (-6.5) µm; hilum 1.5-1.75 µm thick.

On living leaves of Broussonetia papyrifera (L.) Vent.

Coloniae amphigenae, plerumque hypophyllae, raro in petiolis, effusae et saepe pilos folii adscendentes. Mycelium ex hyphis superficialibus, pallidis, ramosis et anastomosantibus, laevibus vel verruculosis, septatis, 1.5-2.5 µm crassis; hyphae sub conidiophoris ad 2.5-6 µm inflatae, brunneae, saepe stroma multicellulare, atro-brunneum, in folii pilis formantes. Conidiophora apicalia et lateralia, macronematosa, mononematosa, solitaria. Stipites erecti, recti vel flexuosi, laeves vel verruculosi, praecipue basi, septati, atro-brunnei, apicem versus pallidiores, usque ad 425 µm longi x 5-11 µm crassi. Rami primarii interdum ramulosi, brunnei, flexuosi, usque ad 75 µm longi x 3.5-5.5 µm crassi. Cellulae conidiogenae polyblasticae, in conidiophoris incorporatae, terminales et intercalares, sympodiales, cicatrices conidiales 1-1.75 µm latae. Conidia solitaria, acropleurogena, pallide brunnea, laevia, 3-4 (-6) septata, constricta vel non constricta, recta vel parum curvata, obclavato-cylindrica, (23-) 25-40 (-51) µm longa x 4.5-5 (-6.5) µm later; cellula apicalis rotundata, cellula basalis obconice truncata; hilo 1.5-1.75 µm lato. In foliis moraceae speciei Broussonetia papyrifera (L.) Vent. Holotypus PDD 50789.

Sporulation may be, but is not necessarily, associated with reddish-brown leaf speckling. The specimen is infected with rust, Cerotelium fici (Cast.) Arthur.

Specimen examined: Solomon Islands, Guadalcanal, Mt Austen, 23.111. 1986, E.H.C. McKenzie (PDD 50789 - holotype).

Specimens examined: New Zealand, Northland, Waima State Forest, Waiotemarama Track, 21.X.1987, E.H.C. McKenzie, P.R. Johnston (PDD 48295); Coromandel, Kauaeranga Valley, Moss Creek Hut Track, 30.VIII.1986, E.H. C. McKenzie (PDD 48296 - holotype); Waiomu, Crosbies Hut Track, 31.VIII. 1986, E.H.C.McKenzie (PDD 48297); Auckland, Waitakere Ranges, Matuku Forest & Bird Reserve, 27.XII.1986, E.H.C. McKenzie (PDD 48298).

Colonies on the adaxial leaf surface. Mycelium superficial and immersed, pale, branched and anastomosing, smooth or verruculose, septate,1.5-3 µm thick. Stroma absent. Conidiophores arising terminally and laterally on the hyphae, macronematous, single, composed of a mononematous stipe and a head of branches bearing conidia, sometimes unbranched. Stipes erect, straight or flexuous, smooth, septate, brown, paler towards the apex, up to 250 µm long x 3-5.5 µm thick at base tapering to 2.5-3.5 µm thick at the apex. Primary branches sometimes bearing branchlets, pale brown, flexuous, up to 100 µm long, usually 15-30 µm long x 1.5-3 µm thick. Conidiogenous cells polyblastic, integrated, terminal, sympodial, conidial scars c. 0.5 µm wide. Conidia solitary, acropleurogenous, pale, verruculose, nonseptate, ellipsoidal to obovoid, apex obtuse, base truncate, (3-) 4.5-6 (-7) x (1.5-) 2-3 µm; hilum c. 0.5 µm thick.

On living leaves of Cordyline banksii Hook. f.

Coloniae adaxiales. Mycelium partim intermum et extermum, ex hyphis pallidis, ramosis et anastomosantibus, laevibus vel verruculosss, septatis, 1.5-3 µm crassis. Stroma nullum. Conidiophora apicalia et lateralia, macronematosa, mononematosa, solitaria. Stipites erecti, recti vel flexuosi, laeves, septati, brunnei, apicem versus pallidiora, usque ad 250 µm longi x 2.5-5.5 µm crassi. Rami primarii interdum ramulosi, pallide brunnei, flexuosi, usque ad 100 µm longi, plerumque 15-30 µm longi x 1.5-3 µm crassi. Cellulae conidiogenae polyblasticae, in conidiophoris incorporatae, terminales, sympodiales, cicatrices conidiales c. 0.5 µm ratae. Conidia solitaria, acropleurogena, pallide, verruculosa, eseptata, ellipsoidea vel obovoidea, (3-) 4.5-6 (-7) x (1.5-) 2-3 µm; hilo c. 0.5 µm lato. In foliis agavaceae speciei Cordyline banksii Hook. f. Holotypus PDD 48296.

Sporulation is usually associated with an indistinct reddish-brown leaf speckle.

Specimens examined: Australia, Queensland, Mt Tamborine, 13.111.1984, J.L. Alcorn (PDD 48819, BRIP 14286); near Mary Cairncross Reserve, 21.111.1988, E.H.C. McKenzie (PDD 53907). Palau, Koror, 6.VI.1986, G.V.H. Jackson (PDD 45618). Vanuatu, Efate, Erakor Village, 21.IV.1983, E.H.C. McKenzie (PDD 46997); same details, 25.II.1985, E.H.C. McKenzie (PDD 46819, 46865 - holotype); Efate, Snake Hill, 21.X1.1983, E.H.C. McKenzie (PDD 46916); Malekula, Brenwei, 23.X1.1983, E.H.C. McKenzie (PDD 46996); Santo, near Hog Harbour, 15.11.1985, E.H.C.McKenzie (PDD46820); Tanna, White Grass, 19.11.1985, E.H.C. McKenzie (PDD 46878).

Colonies hypophyllous, occasionally on petioles and ascending leaf hairs. Mycelium superficial, pale, branched and anastomosing, smooth or verruculose, septate, 1.75-2.5 µm thick. Stroma absent. Conidiophores arising terminally and laterally on the hyphae, macronematous, single, composed of a macrnematous stipe and a complex head of branches bearing conidia. Stipes erect, straight or flexuous, smooth, occasionally verruculose towards apex, septate, brown, paler towards the apex, up to 460 µm long x (3-) 4.5-6 (-6.5) µm thick at base tapering to 2.5-4 µm thick at the apex. Primary branches sometimes bearing branchlets, pale brown, flexuous, up to 70 µm long, usually 15-40 µm long x 2-3 µm thick. Conidiogenous cells polyblastic, integrated, occasionally borne directly on hyphae, terminal, sympodial, conidial scars prominent, 0.75-1 µm wide. Conidia solitary, acropleurogenous, straw-coloured, smooth, non-septate, cylindrical to obovoid, apex obtuse, base truncate, 5-6 (-6.5) x 1.75-2 (-2.5) µm; hilum 0.75-1 µm thick.

On living leaves of Persea americana Mill.

Coloniae hypophyllae, interdum in petiolis, interdum pilos folii adscendentes. Mycelium ex hyphis superficialibus, pallidis, ramosis et anastomosantibus, laevibus vel verruculosis, septatis,1.75-2.5 µm crassis. Stroma nullum. Conidiophora apicalia et lateralia, macronematosa, mononematosa, solitaria. Stipites erecti, recti vel flexuosi, laeves vel saepe apicem versus verruculosa, septati, brunnei, apicem versus pallidiora, usque ad 460 µm longi x 2.5-6.5 µm crassi. Rami primarii interdum ramulosi, pallide brunnei, flexuoui, usque ad 70 µm longi, plerumque 15-40 µm longi x 2-3 µm crassi. Cellulae conidiogenae polyblasticae, in conidiophores incorporatae, terminales, sympodiales, cicatrices conidiales 0.75-1 µm latae. Conidia solitaria, acropleurogena, straminea, laevia, aseptata, cylindrica vel obovoidea, 5-6 (-6.5) x 1.75-2 (-2.5) µm; hilo 0.75-1 µm lato. In foliis lauraceae speciei Persea americana Mill. Holotypus PDD 46865.

Black mould growth is sometimes visible on the undersurface of the leaves. The leaves may show reddish-brown blotches, but fungal sporulation is not restricted to the discoloured areas. Conidiophores are concentrated along the mid-rib and veins.