ADDITIONAL SPECIMENS EXAMINED: ARGENTINA: NEUQUÉN: Parque Nacional Nahuel Huapí, Villa La Angostura, on N. dombeyi, I. J. Gamundí & C. Brion, 1 Jun 1997, BCRU 1766, PDD 70296. RÍO NEGRO: Parque Nacional Nahuel Huapí, L. Nahuel Huapí, Brazo Tristeza, on N. dombeyi, I. J. Gamundí, 3 May 1994, BCRU 1752, PDD 70297; Parque Nacional Nahuel Huapí, path from Puerto Blest to Los Cántaros, on N. dombeyi, I. J. Gamundí, 28 Feb 1994, BCRU 1755; Parque Nacional Nahuel Huapí, Lago Frias, path to Paso de las Nubes, on N. dombeyi, I. J. Gamundí & C. Pujals, 24 Apr 1958, BAFC 20267a; Lago Mascardi, on N. dombeyi, I. J. Gamundí & C. Pujals, 28 Apr 1958, BAFC 20347.

Apothecia developing on both sides of fallen leaves; erumpent through host tissue, arising from small patch of stromatic tissue at base of stipe; apothecia 0.45-1 mm diam., stipitate, fleshy, fragile; disc plane, pale grey to brownish; receptacle grey to pale yellowish brown with scattered, pale, stiff setae; stipe 0.4-1 x 0.15-0.25 mm, narrow-cylindric, concolorous with receptacle, darkened near base, the colour coming from the inner layer. Paraphyses 2-2.5 µm diam., undifferentiated or branched, about same length as asci. Asci 70-106 x 7.5-12 µm, cylindric-clavate, tapering to truncate apex, wall thickened at apex, pore with weak amyloid reaction, 8-spored. Ascospores 16.5-20.5 x 4-5 µm, fusoid, flattened one side, slightly curved, acute to the ends, 2-4 guttulate, hyaline; 1-2-seriate. Ectal excipulum 3-layered; outer layer very poorly developed, lacking on stipe, on receptacle comprising meandering, long-cylindric cells 3-4 µm diam. with walls encrusted, hyaline to pale brown; central layer 30-40 µm wide in receptacle, here comprising broad cylindric cells 15-30 x 6-12 µm with walls slightly thickened, agglutinated, hyaline, in stipe this layer narrower, comprising long cylindric cells 20-40 x 3-6 µm with walls gelatinised, hyaline; inner layer poorly developed in receptacle, in stipe comprising long cylindric cells with walls thinly encrusted, hyaline to pale brown, nongelatinous. Medullary excipulum comprising textura intricata of hyphae 2-5 µm diam. with walls hyaline, nongelatinous. Setae arising from outer excipular layer, 100-250 x 6.5-7.5 µm, slightly swollen near the base, tapering to simple base, tapering gradually toward apex, with upper 2-3 cells swollen to 10-13 µm diam. at the broadly rounded apex, walls smooth, pale brown, upper 2-3 cells hyaline, pluriseptate; setae on stipe slightly shorter, 40-120 µm long.

Apothecia 0.45-1 mm diam., ad folia dejecta, amphigena, erumpentia ex hospiti epidermia, stipitata, margine integerrima; hymenium convexum, griseum vel avellaneum; receptaculum sepiaceum vel umbrinum, parce setosum; stipes 0.4-1 x 0.15-0.25 mm cylindricus, umbrinus ad basim fuligineus, setosus. Paraphyses simplices vel bifurcatae, filiformes, 2-2.5 µm diam. Asci 8-spori, 70-106 x 7.5-12 µm, cylindrico-claviformes, poro iodo coerulescente vel leniter coerulescentes. Ascosporae 16.5-20.5 x 4-5 µm, fusoideae, asymmetricae, 2-4 guttulatae, hyalinae; 1-2-seriatae. Excipulum ectale tristratosum in receptaculo, cum strato externo infirmo evoluto; strato medio hyalino, textura prismatica sub-gelatinosa; strato interno textura porrecta, non gelatinoso, brunnescente. Excipulum ectale bistratosum in stipite, strato externo absente. Excipulum medullare textura intricata, hyalina. Setae ad receptaculum 100-250 x 10-13 µm, ad stipitem breviores (40-120 µm longae), robustae, pluriseptatae, brunneae ad apicem pallidiores, bulbosae vel spathuliformia.

ETYMOLOGY: The specific epithet refers to the pale grey colour of the receptacle and hymenium.

T. grisea, known only from Nothofagus dombeyi, is characterised by the pale brown setae that are swollen and hyaline at the apex. Although unusual for Torrendiella, at least one other species, an undescribed species from Epacridaceae from Australia and New Zealand, has the same kind of setae (P.R. Johnston, unpubl. data). The setae of these fungi are very like those of Zoellneria rosarum, the type of Zoellneria; however that genus differs from Torrendiella in other respects (see Introduction). The 3-layered nature of the ectal excipulum is poorly developed in T. grisea.

Note that the collections BAFC 20267a and 20347 were recorded erroneously by Gamundí (1962) as Zoellneria eucalypti.

Chile: Los Lagos: Parque Nacional Puyehue, road to Hotel Antillanca, on Nothofagus dombeyi, P. R. Johnston SA265 & I. J. Gamundí, 10 Apr 1996, BCRU 3833 - holotype, PDD 70145 - isotype.