Rhizome short, erect, bearing dark brown subulate scales up to 15 × 2 mm. Stipes 5-15 cm long, brown below, green above, covered in subulate to narrowly triangular scales with long filiform apices (Fig. 24J-N, Q-V). Laminae lanceolate to elliptic, 10-30 × 5-20 cm, dark green, leathery or very thick and fleshy, slightly drooping or stiff and upright, bi- to tripinnate. Raches green, scaly, prominently ridged. Pinnae 7-20 pairs, those at the base ovate, those above very narrowly ovate or elliptic, acute to acuminate, stalked, 3-10 × 0.5-3 cm, covered on underside in tiny scales. Upper pinnules linear, acute, entire and sessile; lower pinnules ovate to narrowly elliptic and pinnatifid or pinnate, up to 30 × 10 mm. Sori submarginal, 2-7 mm long. Spores (38)43-51(56) mcm long, (25)29-34(38) mcm wide (14 populations), perispore prominently winged and ridged (Fig. 30 O-R). 288 chromosomes in the sporophyte; 144 bivalents at meiosis.

Rhizoma breve, erectum, paleis fuscis subulatis usque ad 15 × 2 mm obtectum. Stipes 5—15 cm longus, infra fuscus, supra viridis, paleis subulatis vel anguste triangularibus cum apicibus longis filiformibus vestitus (Fig. 24J-N, Q-V). Lamina lanceolata vel elliptica, 10-30 × 5-20 cm, atroviridis, coriacea aut crassissima carnosaque, cernua aut rigida erectaque, bi- vel tri-pinnata. Rhachis viridis, squamata, manifeste canaliculata. Pinnae 7-20 jugatae, inferiores ovatae, superiores peranguste ovatae vel ellipticae, acutae vel acuminatae, 3-10 × 0.5-3 cm, paleis minutis infra vestitae. Pinnulae superiores lineares acutae integrae sessilesque; inferiores ovatae vel anguste ellipticae, pinnatifidae vel pinnatae, petiolatae, usque ad 30 X 10 mm. Sori submarginales, 2-7 mm longi. Sporae (38)43-51(56) mcm longae, (25)29-34(38) mcm latae, perisporae manifeste alatae cristataeque (Fig. 30 O-R). Numerus somaticus chromosomatum 288; in meiosi paria 144 observantur.