UNKNOWN HOSTS. Queensland, Dunk Island (herb. Kew); Toowoomba (herb. Kew); Moreton Bay (herb. Kew). New South Wales, Shoalhaven Rivet (herb. Kew); Kendall; Dorrigo; Lisarow; Bullahdelah; Bulli; Kangaroo Valley. Norfolk Island.

Hymenophore annual, coriaceous, centrally stipitate. Pilei commonly infundibuliform, sometimes with one side partly suppressed, or 2-3 flabelliform pilei may merge to form rosetted clusters, 6-12 x 4-10 cm; pileus surface cream or wood colour, coarsely and radiately ridged, acute margins sometimes toothed, with between ridges a dense tomentum to 3 mm thick and lighter in colour; margins acute, plane or slightly inrolled, crenate and torn; hymenial surface decurrent, cream or buff, with radiate rounded ridges, which may be branched towards apices, bearing a few scattered papillae. Stems 3-5 cm tall, 5-15 mm diameter, occasionally shorter and often merged at the base into several, usually clothed with dense tomentum with a darker margin where the hymenium ends, attached by a prominent mycelial disc. Context white, to 1 mm thick, becoming progressively thinner from base to apex, of radiately arranged parallel hyphae, bordered by a cemented cortex of hyaline intertwined hyphae bearing abhymenial hairs; skeletal hyphae 3-4.5 µmdiameter, lumena almost capillary; generative hyphae 3-4 µm diameter, walls 0.2 µm thick, with clamp connections. Gloeocystidia either fusiform or somewhat lanceolate, arising from the subhymenium but scarcely projecting, when 32-60 x 6-12 µm, with thin walls, subglobose or oval when to 12 µm diameter or (as 'cystidia') broadly pyriform to conical, with inflated bases and acute apices, 32-48 x 12-24 µm, walls thickened to 3 µm, upper parts encrusted with coarse deciduous crystals. Hymenial layer to 130 µm deep, a dense palisade of basidia, paraphyses, gloeocystidia, and 'cystidia'. Basidia subclavate, 24-30 x 6-7 µm, bearing 4 spores; sterigmata erect, stout, to 5 µm long. Paraphyses subclavate, 18-22 x 6-7 µm. Spores broadly elliptical, 6-7.5 x 4-4.5 µm, walls smooth, hyaline, 0.1 µm thick.

DISTRIBUTION: Fiji, Australia, Norfolk Island.

HABITAT: Solitary or caespitose on bark of dead branches.

From S. caperatum the species may be separated by one feature alone, the presence of cystidial in the-hymenial layer. These organs differ markedly from metuloids of Peniophora for they are without pedicels, of different shape, and crystals which encrust parts are deciduous and disappear in aqueous-solutions of potassium hydroxide. Their appearance and the presence of many transitional stages between them and gloeocystidia indicate that they are old gloeocystidia in which walls have become thickened and partly covered with crystals. All stages between them and typical gloeocystidia may be seen in sections, and they are present also in rudimentary form in S. caperatum; although without crystals in that species. Study of a large series of specimens will no doubt show that the feature is not constant and that plants are merely forms of one variable species.
Pilei vary appreciably in shape, size, length of stems, and in surface features. Commonly infundibuliform, specimens examined may consist of two or several fused either at the base or margins; several pilei may become flabelliform then fuse at margins to form rosettes; or pilei may split on one side and only partly develop. One stem may support two pilei 1 or conversely several stems may develop but fuse in rosetted plants. Stems may be long and cylindrical, or short and tapering, be round, flattened, or often fluted, and are usually clothed, with coarse tomentum, although sometimes naked. The surface may bear few or many radiate lamellae with acuminate margins, their number, size, and colour varying in almost every -specimen examined: Between them develops a dense tomentum which maybe 3 mm deep, and vary in colour from cream to buff; or be suppressed, when the naked pileus appears reddish-brown.

TYPE LOCALITY. Fiji: