Garnock-Jones, P.J.; Bayly, M.J.; Lee, W.G.; Rance, B.D. 2000: Hebe arganthera (Scrophulariaceae), a new species from calcareous outcrops in Fiordland, New Zealand. New Zealand Journal of Botany 38(3): 379–388.
Details
Details
Garnock-Jones, P.J.; Bayly, M.J.; Lee, W.G.; Rance, B.D. 2000: Hebe arganthera (Scrophulariaceae), a new species from calcareous outcrops in Fiordland, New Zealand. New Zealand Journal of Botany 38(3): 379–388.
Article
Descriptions
Shrub, often rounded or spreading, to 0.5 m tall. Branches ascending to erect; old stems brown; youngest branchlets green; internodes (2-)4-8 mm long, bifariously eglandular-pubescent; leaf decurrencies evident; leaf-base scars prominent. Leaf bud about as long as mature leaves, with leaves of a pair separating when mature; sinus narrow, acute. Leaves decussate, free at base, erecto-patent to patent; lamina oblong to elliptic, coriaceous or subcoriaceous, m-shaped in transverse section, ( 12-) 15-30(-38) × (5-)6- 1 1 mm; apex subacute and mucronate; base cuneate; midrib thickened beneath and depressed to grooved above; secondary veins not evident; margin entire, translucent, bevelled or rounded, minutely papillate or denticulate; adaxial surface green or yellowish green, dull, with many stomata, hairy along midrib; abaxial surface pale green, dull, densely covered with stomata, glabrous. Petiole 3-4 mm long, uniformly eglandular-pubes- cent. Inflorescences with 15-20(-25) flowers, lateral, racemose, simple or with (1-)2 branches at base, (2-)4-5 cm long, about = or > subtending leaves, flowers opening in acropetal sequence, usually all developing to maturity; peducle 0.8-1 cm long, eglandular-pubescent; rachis 1. 5 4 cm long, eglandular pubescent; bracts opposite and decussate at least below, sometimes becoming alternate above, subacute to acute, eglandular-ciliolate, linear to lan- ceolate; pedicels < bracts, eglandular-pubescent, erecto-patent at flowering and fruiting, (0.5-) 1.5-3(- 5) mm long. Flowers protandrous, all hermaphrodite. Calyx rounded at base, terete, 4-lobed, equally di- vided, 2.5-3.5 mm long; lobes all similar, lanceolate to elliptic, obtuse, eglandular-ciliolate; margins scarious. Corolla white at anthesis and after polli- nation; tube glabrous, 1-2 mm long, c. 1 mm wide, cylindric, > calyx; lobes glabrous, > tube; posterior lobe circular to elliptic, obtuse, erecto-patent; lateral lobes elliptic or ovate, obtuse, erecto-patent, cuneate at base; anterior lobe ovate, obtuse, patent to recurved, not enfolding style; corolla throat white. Stamens: filaments white, diverging after anthesis, straight at apex in bud, (4-)5-6 mm long; anthers subacute to acute, white, c. 1.5 mm long. Nectarial disc glabrous. Ovary ovoid, glabrous, 1.7-2 mm long, bilocular; ovules 20-25 per locule, marginal in 1-2 layers on a flattened placenta; style 5-7 mm long, c. 0.2 mm thick (when dry), glabrous, white; stigma capitate, 0.3-0.4 mm wide. Capsules latiseptate, acute, dark brown, 3-4 × 2.5-3.5 mm, 1.5-2.5 mm thick, glabrous; septicidal split extend- ing to base, loculicidal split extending ¼-way to 1/3- way to base; capsule valves with pronounced midrib. Seeds 6- 10 (or more) per locule, flattened, ellipsoid, not winged, smooth, straw yellow to pale brown, 0.6-1 × 0.5-0.8 mm.
Hebes cockayneanae et H. rupicolae similis, sinu angusto, cicatricibus in caulibus prominentibus, capsulis brevibus latis; a H. cockayneana pubescentia caulis breviore, foliis grandioribus viridibus concoloris, stomatibus utrinque manifestis, marginibus minute eglandulato-ciliolatis, floribus plerumque oppositis, calyce eglandulato-ciliolatis, seminibus parvioribus differt; a H. rupicola cicatricibus in caulibus prominentioribus, foliis utrinque viridibus, inflorescentiis hirsutioribus, calyce eglandulato-ciliolatis, lobis corollae obtusatis, capsulis latioribus pallidioribus, seminibus parvioribus differt.
Identification keys
KEY TO THE SPECIES OF HEBE IN FIORDLAND NATIONAL PARK
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Inflorescences terminal (but often also with some lateral inflorescences), fertile shoots not terminated by a vegetative bud
Inflorescences strictly lateral (but possibly in axils of uppermost leaves, and therefore appearing terminal without close inspection), fertile shoots terminated by a vegetative bud
4
Leaves dark green and glossy above, paler below; leaf bases free; vegetative branches terminated by a prominent leaf bud with a shield-shaped sinus at its base (Fig. 5P); rounded shrub of grassland, bog margins and scrub
Leaves dull yellowish-green above and beneath; leaf bases of each leaf pair shortly connate (Fig. 5F, G); vegetative branches not terminated by a prominent leaf bud (i.e., apical leaves not generally cohering along their margins); low-growing subshrub of exposed rocks and scree
Leaves usually > 10 mm long (Fig. 5E, L), usually glossy, never glaucous; flowers pedicillate
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Midrib forming a rounded keel on underside of leaf (Fig. 5Q, cf. Fig. 5S); leaf margin fringed with fine, white, tangled, sometimes branched hairs (Fig. 5R); anterior calyx lobes fiee
Keel of leaf usually characteristically Battened toward apex (Fig. 5S); leaf margin glabrous or with minute caducous cilia, anterior calyx lobes usually fused below emarginate tip
10
Leaf margin white pubescent except near strongly mucronate apex (Fig. 5D) and well-defined petiole; shrub of coastal rocks
Leaf margin glabrous or minutely or sparsely hairy, without such a strong contrast of indumentum at petiole and apex; shrubs of upland areas
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Leaf decurrencies prominently swollen; leaf bud sinus broad, shaped somewhat like a vaulted arch (Fig. 5V); leaf margins minutely crenulate (Fig. 5W)
Leaf decurrencies not prominently swollen; leaf bud sinus comparatively narrow and acute (e.g., Fig. 5K); leaf margins entire, minutely papillose on lower surface (Fig. 5N) or cartilaginous
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Leaves dull green above, glaucous below (except along margins and midrib), (5) 10-22 mm long; leaf margins glabrous (apart from minute papillae, Fig. 5N), or with scattered glandular hairs (Fig. 4B)
Leaves dull green, f concolorous, (12-)15-30(-38) mm long; leaf margins ciliate with many minute eglandular hairs (Fig. 4A)
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Leaves ribbed or striped on outer surface, at least for a short distance from upper margin (Fig. 5H)
Reproduced with permission from New Zealand Journal of Botany and The Royal Society of New Zealand Te Apārangi.
Cited scientific names
Metadata
bfddc986-0d74-4193-92f9-16348dd12287
reference
Names_Plants
18 October 2022
